Crataegus berberifolia |
Crataegus pennsylvanica |
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barberry hawthorn, barberry-leaf hawthorn |
Pennsylvania hawthorn, Pennsylvania thorn |
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Habit | Shrubs or trees, 60 dm. | Shrubs or trees, 70–80 dm. | ||||
Stems | twigs: new growth orange-brown or green tinged with red, ± pubescent, 1-year old brown, older gray; thorns on twigs ± straight or recurved, 2-years old shiny black to chestnut brown, fine or stouter, (2–)3–4(–6.5) cm. |
trunk bark newly exposed orange-brown, older buff, fibrous, checked into longitudinal plates; twigs: new growth densely pubescent, 1-year old fawn, older gray; thorns on twigs absent or few, usually recurved, 2-years old shiny dark brown to blackish, stout to more slender, 3–5 cm. |
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Leaves | petiole 4–6 mm, length 13–18% blade, glabrescent, eglandular; blade narrowly obovate to oblanceolate, (2.5–)3(–4) cm, coriaceous, base narrowly cuneate, lobes 0, margins finely crenate or serrate except at base, or only beyond widest part, venation craspedodromous, veins 4–6 per side, apex subacute to obtuse, lustrous, abaxial surface ± densely pilose on veins, sometimes pubescent on surface, adaxial hairy young, glabrescent or becoming scabrous. |
stipules axillary, circinate, herbaceous, gland-bordered; petiole relatively slender, stouter than C. coccinea, length 30–35% blade, densely pubescent young, sparsely mature, often sparsely stipitate-glandular; blade broadly ovate to ovate-deltate, 6–10 cm, slightly coriaceous, base broadly cuneate to truncate, lobes 4–6 per side, sinuses shallow, lobe apex acute to obscure, margins sharply serrate, teeth at base often stipitate-glandular, veins 5–7 per side, apex acute, abaxial surface sparsely scabrous-pubescent young, glabrescent, veins hairy mature, adaxial densely appressed-scabrous young. |
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Inflorescences | 8–12-flowered; branches densely pubescent; bracteoles linear, margins glandular. |
8–20-flowered; branches tomentose; bracteoles variably persistent, linear-lanceolate, membranous to (larger) ± herbaceous, margins glandular. |
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Flowers | 10–20 mm diam.; hypanthium villous or glabrous; sepals 3–5 mm, margins entire, abaxially glabrous; stamens 10 or 20, anthers cream or pink; styles 2 or 3. |
17–21 mm diam.; hypanthium densely tomentose; sepals narrowly triangular, 7–8 mm, margins glandular-laciniate, abaxially glabrate; stamens 5 or 6, or 10, anthers cream to pale pink or salmon; styles 4 or 5. |
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Pomes | reddish to yellow, suborbicular, 8–10 mm diam., glabrous; sepals erose or patent; pyrenes 2 or 3. |
bright red, suborbicular, 10–12 mm diam., densely villous; sepals prominent, erect-patent; pyrenes 4 or 5. |
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Crataegus berberifolia |
Crataegus pennsylvanica |
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Phenology | Flowering May–Jun; fruiting Sep–Oct. | |||||
Habitat | Brush, successional fields, open woodlands, fencerows | |||||
Elevation | 10–200 m (0–700 ft) | |||||
Distribution |
AL; AR; FL; GA; KS; LA; MO; MS; NC; SC; TN
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DE; NC; NY; OH; PA; ON |
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Discussion | Varieties 2 (2 in the flora). Crataegus berberifolia is widespread from Texas to Missouri, Florida, and Virginia; it is particularly abundant in Louisiana. Crataegus berberifolia is little differentiated from some forms of C. crus-galli, except in indumentum; it has relatively small and less variably shaped leaves. Its distribution is quite different. Intermediates with hairy leaves and glabrous inflorescences or nearly glabrous leaves and hairy inflorescences may represent hybrids with C. crus-galli (where they might be reached in the key). Such a situation is found in C. araioclada. Abrasion of the adaxial leaf pubescence may occur, rendering identification more difficult with fruiting material. Crataegus berberifolia has a plethora of yellow and orange-fruited forms, particularly from southern Louisiana. Crataegus fera and C. tersa are red-fruited, C. crocina yellow. The fruit color in the type is unknown; E. J. Palmer (in specimen annotation) called it yellow-orange. A form with exceptionally white-tomentose leaves near Copenhagen, Louisiana, is probably this species. Crataegus regalis var. paradoxa (Sargent) E. J. Palmer, from Missouri and adjacent Kansas and Arkansas, is probably a hybrid between the deeply serrated 'regalis' leaf form of C. crus-galli and a form of C. berberifolia. Two common forms of C. berberifolia occur, treated here as varieties: var. engelmannii with ten pink anthers and var. berberifolia with 20 cream anthers. Forms with 20 pink or ten cream anthers also occur sporadically. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Crataegus pennsylvanica ranges mainly to the south of C. submollis and may be locally common. A highly disjunct record from North Carolina (Buncombe County) appears unequivocal. Crataegus pennsylvanica has been confused with C. mollis and various members of ser. Coccineae. Except for a small overlap in northeastern Ohio, its range lies outside that of C. mollis. Crataegus pennsylvanica is more similar to C. submollis, from which it differs by its less variable leaves (usually broadly ovate, more or less truncate, relatively large), by its much less thorny nature, by its indumentum on young parts being tomentose rather than densely pubescent, and by its being more treelike (C. submollis is usually a shrub). Thorns are absent or sparse on older trees. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 9, p. 540. | FNA vol. 9, p. 557. | ||||
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Crus-galli | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Crataegus > sect. Coccineae > ser. Molles | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | C. tatnalliana | |||||
Name authority | Torrey & A. Gray: Fl. N. Amer. 1: 469. (1840) | Ashe: Ann. Carnegie Mus. 1: 394. (1902) | ||||
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