Cotula |
Cotula coronopifolia |
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cotula, water-buttons |
brass-buttons, buttonweed, common brass-buttons, cotule, golden buttons |
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Habit | Annuals or perennials, 2–25[–50+] cm (sometimes aromatic). | Perennials, (3–)5–15(–25+) cm tall or across. | ||||
Stems | usually 1, erect or prostrate to decumbent or ascending (sometimes rooting at nodes), usually branched, glabrous or ± strigillose to villous (hairs mostly basifixed). |
prostrate (rooting at nodes, ± fleshy) to decumbent or erect, glabrous. |
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Leaves | usually mostly cauline [basal]; alternate [opposite]; petiolate or sessile; blades obovate or spatulate to lanceolate or linear, sometimes 1–3-pinnately [palmati-pinnately] lobed, ultimate margins entire or irregularly toothed, faces glabrous or ± strigillose to villous [lanate] (hairs mostly basifixed). |
sessile (bases sheathing stems); blades linear to lanceolate or oblong (sometimes lobed, lobes lanceolate to linear), (1–)2–3(–7) cm, ultimate margins entire or irregularly toothed or lobed, faces gland-dotted. |
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Involucres | broadly hemispheric to saucer-shaped, 3–12+[–15+] mm diam. |
6–9(–12+) mm diam. |
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Receptacles | flat to convex [conic], epaleate (sometimes ± covered with persistent stalks of florets). |
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Ray florets | 0 [5–8+, pistillate, fertile; corollas white] (peripheral pistillate florets 8–80+ in 1–3+ series; corollas usually none). |
0 (peripheral pistillate florets 12–40+ in 1 series; corollas 0). |
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Disc florets/ |
12–200+[–600+], bisexual, fertile [functionally staminate]; corollas ochroleucous or yellow, tubes ± cylindric (bases sometimes adaxially saccate), throats abruptly ampliate, lobes (3–)4, ± deltate (sometimes one larger than others, usually each with central resin canal). |
yellow, 1–1.5 mm. |
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Phyllaries | persistent, 13–30+ in 2–3+ series, margins and apices (colorless, light to dark brown, or purplish) scarious. |
21–30+ in 2–3+ series. |
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Heads | disciform [discoid or radiate], borne singly (peduncles sometimes dilated). |
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Cypselae | obovoid to oblong, ob-compressed or -flattened, ribs 2, lateral, sometimes becoming wings, faces ± papillate (pericarps relatively thin, sometimes with myxogenic cells and/or 2 lateral resin sacs); pappi 0. |
outer 1.2–1.8 mm, winged, adaxial faces papillate; inner 0.7–1 mm, scarcely, if at all, winged, not papillate. |
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x | = 10. |
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2n | = 20. |
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Cotula |
Cotula coronopifolia |
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Phenology | Flowering Mar–Dec. | |||||
Habitat | Saline and freshwater marshes, along streams | |||||
Elevation | 0–900 m (0–3000 ft) | |||||
Distribution |
s Old World [Introduced in North America; introduced also (perhaps some native) in Mexico, South America, s Oceanic Islands] |
AK; AZ; CA; MA; MD; NV; OR; WA; BC; NB; NS; PE; QC; South America; S Africa; Australia; Europe [Presumed introduced; also introduced in Mexico (Baja California)]
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Discussion | Species 55 (2 in the flora). Some species of Cotula are widely naturalized. F. Hrusa et al. (2002) reported Cotula mexicana (de Candolle) Cabrera as established on golf courses in California; it is similar to C. australis and differs in leaf blades mostly 1-pinnate, receptacles pilose, and disc florets functionally staminate. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 543. | FNA vol. 19, p. 544. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Name authority | Linnaeus: Sp. Pl. 2: 891. (1753): Gen. Pl. ed. 5, 380. (1754) | Linnaeus: Sp. Pl. 2: 892. (1753) | ||||
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