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Mclaren's cotoneaster

Himalayan cotoneaster, khasiberry cotoneaster, simons' cotoneaster

Habit Shrubs or trees, 4–6 m. Stems loosely erect, spreading, stiff; branches spiraled, maroon, minutely verrucose, initially pilose-strigose. Shrubs, 2–4(–6) m; ± strict or ascending, crown narrow or ± columnar.
Stems

erect, stiff;

branches spiraled, divaricate, lateral branches ± straight, mostly suppressed, short, grayish brown, initially densely strigose.

Leaves

deciduous;

petiole 4–6 mm, densely pilose-strigose;

blade on sterile shoots elliptic, (35–)50–80(–103) x (13–)17–37(–42) mm, chartaceous, margins flat, base cuneate, veins 6–10, superficial or lightly sunken, apex acute or obtuse, abaxial surfaces color not recorded, villose-strigose, adaxial brownish green, soon green, dull, coating not recorded, flat between lateral veins, glabrate;

fall leaves yellow.

deciduous (sometimes tardily so on vigorous sprouts);

petiole 3–5 mm, strigose;

blade on vigorous shoots broadly elliptic to ovate, sometimes elliptic, rarely suborbiculate, 17–36 × 10–25 mm, chartaceous or subcoriaceous, base obtuse or cuneate, margins flat, veins 4 or 5, superficial or slightly sunken, apex acuminate or acute, abaxial surfaces light green, initially strigose, usually becoming sparsely so, adaxial green to dark green, shiny, not glaucous, flat or faintly bulging between lateral veins, or slightly rugose, sparsely strigose;

fall leaves yellow to bright orange or fiery red.

Inflorescences

on fertile shoots 40–50 mm with 2–4 leaves, 10–25-flowered, lax.

on fertile shoots 15–30 mm with 4 leaves, 2–6-flowered, compact.

Pedicels

3–5 mm, densely pilose-strigose.

2–5 mm, strigose.

Flowers

10–12 mm diam.;

buds white;

hypanthium cupulate, villose-strigose;

sepals: margins villous, apex acute, surfaces sparsely villous;

petals spreading, white, with large hair tuft;

stamens (16–)20, filaments white, anthers black, styles 1 or 2.

pendent, (5–)8–10 mm, open;

hypanthium campanulate, strigose;

sepals: margins villous, borders narrow, glabrous, apex recurved, long acuminate or cuspidate, surfaces strigose;

petals erect-incurved, dark red with white border;

stamens 20, filaments pink, anthers white;

styles (2 or)3–5.

Pomes

maroon to purple-black, obovoid to elliptic-obovoid, 9–12 × 9–10 mm, slightly shiny, glaucous, sometimes sparsely villous;

sepals flat, margins glabrous, sparsely villous;

navel slightly open;

style remnants 9/10 from base.

bright orange to orange-red, usually obovoid, rarely cylindric or broadly obovoid, 7–14 × 6–10 mm, shiny, not glaucous, glabrous or sparsely villous;

sepals erect or ascending, rarely nearly flat, margins villous, glabrous or sparsely villous;

navel open;

style remnants 2/3 from base.

Pyrenes

1 or 2.

(2 or)3–5.

Cotoneaster transens

Cotoneaster simonsii

Phenology Flowering Jun; fruiting Oct–Nov. Flowering May–Jun; fruiting Oct–Apr.
Habitat Thickets Shores, clearings, disturbed or open forests, thickets
Elevation 0–50 m (0–200 ft) 0–200 m (0–700 ft)
Distribution
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
CA; OR; TN; WA; BC; Asia (Bhutan, India) [Introduced in North America; introduced also in Europe, Pacific Islands (New Zealand)]
[WildflowerSearch map]
[BONAP county map]
Discussion

L. Lingdi and A. R. Brach (2003) synonymized Cotoneaster transens with C. glaucophyllus Franchet (as C. glaucophyllus var. glaucophyllus). Cotoneaster transens is a deciduous species with purple-black pomes and is not closely related to C. glaucophyllus, an evergreen species with smaller leaves, smaller red pomes, and smaller flowers, among other differences.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Cotoneaster simonsii from Ohio (M. A. Vincent and A. W. Cusick 1998; T. S. Cooperrider et al. 2001) are here referred to C. divaricatus. The binomial C. symondsii T. Moore was proposed as an earlier name for C. simonsii (A. Kumar and G. Panigrahi 1992; W. B. Dickoré and G. Kasperek 2010), but J. Fryer and P. F. Zika (2014) typified C. symondsii so that it is a synonym of the high Himalayan species C. marginatus Lindley ex Loudon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 457. FNA vol. 9, p. 460.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Name authority G. Klotz: Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 17(3): 337, fig. 4. (1968) Baker: in W. W. Saunders, Refug. Bot. 1: plate 55. (1869)
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