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cotoneaster, Tengyueh cotoneaster

Swedish cotoneaster

Habit Shrubs, 2–2.5 m; crown ± rounded. Shrubs, 0.4–0.6 m. Stems ± erect or ± procumbent, arching, ends rooting; branches spiraled, green proximally, purple-black distally, initially densely strigose.
Stems

erect, arching, long;

branches distichous, grayish maroon, initially densely strigose.

Leaves

deciduous (sometimes tardily so on vigorous shoots);

petiole 1–4 mm, strigose;

blade ovate or elliptic, (25–)30–50(–61) x 12–32 mm, chartaceous, base cuneate or rounded, margins flat, veins 3–5, deeply sunken, apex acuminate or acute, abaxial surfaces grayish green, villose-strigose or sparsely to moderately gray-tomentose, adaxial green to dark green, shiny, not glaucous, flat or faintly bulging between lateral veins, pilose-strigose;

fall leaves yellow to orange.

persistent;

petiole often red, 1–3 mm, strigose;

blade elliptic, 10–27 × 4–12 mm, coriaceous, base cuneate, margins revolute, veins 4–7, superficial, apex obtuse or acute, rarely emarginate, abaxial surfaces gray-green, initially strigose-villous, midrib often red, adaxial dark green, shiny, not glaucous, flat between lateral veins, initially sparsely long-haired pilose.

Inflorescences

on fertile shoots 25–40 mm with (3 or)4 leaves, 3–7(–9)-flowered, compact.

on fertile shoots 8–40 mm with 3 or 4 leaves, 1 or 2(–6)-flowered, compact.

Pedicels

1–4 mm, densely strigose.

1–3.5 mm, initially sparsely pilose.

Flowers

erect or ascending, [5–]6–8 mm, opening small;

hypanthium cupulate, strigose;

sepals: margins villous, borders reddish brown, glabrous, apex acuminate or cuspidate, surfaces strigose;

petals erect-incurved, pink to red, base dark pink or red, rarely dark red or maroon, margins white;

stamens 20 or 21, filaments pink or pale pink, whitish distally, anthers white;

styles (2 or)3–5.

12–13 mm diam.;

buds white;

hypanthium cupulate, initially sparsely pilose;

sepals: margins villous, apex acute, surfaces villous;

petals spreading, white, glabrous;

stamens 15–20, filaments white, anthers dark red-purple;

styles 2–4(or 5).

Pomes

spreading or pendent, bright red, broadly obovoid or subglobose, rarely oblong-obovoid, 6.3–10.4 × 5.8–9.1 mm, shiny, not glaucous, sparsely strigose;

sepals flat or suberect, strigose;

navel closed;

style remnants 2/3 from base.

red-orange to scarlet, obovoid to subglobose, [4–]7–11 × 7–10 mm, shiny, not glaucous, glabrous or sparsely villous;

sepals depressed to slightly ascending, glabrous or sparsely villous;

navel open;

style remnants at apex.

Pyrenes

(2 or)3–5.

2–4(or 5) (always some with more than 2).

2n

= 68 (Germany).

= 34 (Germany).

Cotoneaster tengyuehensis

Cotoneaster ×suecicus

Phenology Flowering May–Jun; fruiting Sep–Feb. Flowering May–Jun; fruiting Oct–May.
Habitat Thickets, creek banks, open forests, edges Hedges, edges, paths, urban waste ground
Elevation 0–50 m (0–200 ft) 0–200 m (0–700 ft)
Distribution
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
OR; WA; BC; QC; Europe [Introduced in North America]
[BONAP county map]
Discussion

L. Lingdi and A. R. Brach (2003) confused Cotoneaster tengyuehensis with C. glomerulatus W. W. Smith, a species with small fruits 5 mm in diameter, with 5 pyrenes and styles, and slightly smaller leaves to 40–50 mm. In the flora area, C. tengyuehensis is more likely to be mistaken for C. franchetii, which is a true evergreen with orange-red, obovate to obconic fruits, and pink to purple stamens. Naturalized shrubs of C. tengyuehensis drop their foliage in midwinter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cotoneaster x\suecicus is of garden origin, presumably a cross between C. conspicuus and C. dammeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 465. FNA vol. 9, p. 460.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus
Name authority J. Fryer & B. Hylmö: Watsonia 21: 338. (1997) Cotoneaster ×suecicus G. Klotz: Beitr. Phytotax. 10: 47. (1982)
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