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bullate cotoneaster, puckered-leaf cotoneaster

Stern's cotoneaster

Habit Shrubs, 2–5 m. Stems erect, arching; branches distichous or spiraled, maroon to brown, lenticellate, initially pilose-strigose. Shrubs, 1.5–4 m. Stems erect, arching, stiff; branches distichous, maroon, whitish tomentose-pilose.
Leaves

deciduous;

petiole 0–3 mm, pilose-strigose;

blade elliptic, sometimes ovate-elliptic, rarely obovate, (50–)70–124(–210) x (24–)30–55(–90) mm, subcoriaceous, base cuneate, obtuse, truncate, or auriculate, margins flat, not revolute, veins 8–11, deeply sunken, apex long-acuminate, sometimes acute on short shoots, abaxial surfaces light green, initially yellowish strigose, adaxial green, shiny, not glaucous, strongly wrinkled or bulging between lateral veins (bullate), initially sparsely pilose;

fall leaves intense butter yellow and reddish purple.

persistent;

petiole 4–6 mm, tomentose-pilose;

blade elliptic to broadly ovate, (18–)25–52(–56) x (12–)16–28(–34) mm, coriaceous, base obtuse or cuneate, margins recurved, veins 4–6, deeply sunken, apex acute or acuminate, abaxial surfaces whitish tomentose, adaxial dark green, slightly shiny, not glaucous, on vigorous shoots usually bulging slightly between lateral veins (unless shaded), very rugose, pilose.

Inflorescences

on fertile shoots 60–120 mm with 4 leaves, 8–32-flowered, lax.

on fertile shoots 25–70 mm with 3–5 leaves, 7–20-flowered, compact.

Pedicels

2–4 mm, pilose-strigose.

1–4 mm, tomentose-pilose.

Flowers

(5–)7–9 mm, closed;

hypanthium shallowly cupulate, shiny, sparsely pilose-strigose;

sepals: margins villous, apex acute, surfaces shiny, glabrous;

petals erect-incurved, pink, red, greenish pink, or maroon, margins pink;

stamens 20–23, filaments pink, white distally, anthers white;

styles (4 or)5.

(5–)6–7.5 mm, opening small;

hypanthium cupulate, silky tomentose;

sepals: margins villous, borders purple, glabrous, apex acute or acuminate, surfaces silky tomentose;

petals erect-incurved, pink to red, base dark red (with minute black dots), margins erect-incurved, pink or white;

stamens 20, filaments red, distally pink or whitish, anthers white, sutures pink-tinged;

styles (2 or)3 or 4(or 5).

Pomes

bright to dark red, obovoid, broadly obovoid, or obconic, rarely globose or depressed-globose, 8–12.4 × 7–11.3 mm, shiny, not glaucous, sparsely pilose;

sepals flat, glabrous;

navel closed;

style remnants 3/4 from base.

bright orange, eventually orange-red, subglobose to depressed-globose or globose, 6–10 × 6–10 mm, base rounded, shiny, not glaucous, pilose;

sepals flat, tomentose; forming 5-pointed star over closed navel;

style remnants 3/4 from base.

Pyrenes

(4 or)5.

(2 or)3 or 4(or 5).

Cotoneaster rehderi

Cotoneaster sternianus

Phenology Flowering May–Jun; fruiting Sep–Dec. Flowering Jun–Jul; fruiting Oct–Jan.
Habitat Thickets, disturbed forests, flood plains, lakeshores Mossy cliffs, thickets, hedges, urban waste ground
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AK; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
BC; Asia (China, Myanmar) [Introduced in North America; introduced also in Europe, Pacific Islands (New Zealand)]
[BONAP county map]
Discussion

Reports of Cotoneaster bullatus (treated as C. bullatus var. bullatus by L. Lingdi and A. R. Brach 2003) from British Columbia (J. Pojar 1999) are here referred to C. rehderi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cotoneaster sternianus is similar to C. franchetii, a species with more elongated pomes, uniformly pink or purple anthers, and less textured leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 464. FNA vol. 9, p. 466.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. bullatus var. macrophyllus C. franchetii var. sternianus
Name authority Pojarkova: Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17: 184. (1955) (Turrill) Boom: Jaarb. Ned. Dendrol. Ver. 20: 81. (1957)
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