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bullate cotoneaster, puckered-leaf cotoneaster

silver-leaf cotoneaster, woolly cotoneaster

Habit Shrubs, 2–5 m. Stems erect, arching; branches distichous or spiraled, maroon to brown, lenticellate, initially pilose-strigose. Shrubs, 1–5 m. Stems erect, wide-spreading, arching, pendulous, slender; branches spiraled, purple-black, tomentose-villous.
Leaves

deciduous;

petiole 0–3 mm, pilose-strigose;

blade elliptic, sometimes ovate-elliptic, rarely obovate, (50–)70–124(–210) x (24–)30–55(–90) mm, subcoriaceous, base cuneate, obtuse, truncate, or auriculate, margins flat, not revolute, veins 8–11, deeply sunken, apex long-acuminate, sometimes acute on short shoots, abaxial surfaces light green, initially yellowish strigose, adaxial green, shiny, not glaucous, strongly wrinkled or bulging between lateral veins (bullate), initially sparsely pilose;

fall leaves intense butter yellow and reddish purple.

persistent;

petiole 4–7 mm, tomentose;

blade elliptic, sometimes broadly elliptic, 15–35(–39) x 10–19(–26) mm, mid-coriaceous, base cuneate, margins flat, veins 4–6, usually superficial, sometimes slightly sunken, apex acute or obtuse, abaxial surfaces whitish tomentose, adaxial green to blue-green, dull, not glaucous, usually flat between lateral veins, initially sparsely pilose.

Inflorescences

on fertile shoots 60–120 mm with 4 leaves, 8–32-flowered, lax.

on fertile shoots 20–45 mm, usually with 4 leaves, (3–)5–15(–25)-flowered, compact.

Pedicels

2–4 mm, pilose-strigose.

1–5 mm, tomentose.

Flowers

(5–)7–9 mm, closed;

hypanthium shallowly cupulate, shiny, sparsely pilose-strigose;

sepals: margins villous, apex acute, surfaces shiny, glabrous;

petals erect-incurved, pink, red, greenish pink, or maroon, margins pink;

stamens 20–23, filaments pink, white distally, anthers white;

styles (4 or)5.

[7–]8.5–10 mm diam.;

buds white;

hypanthium funnelform or cupulate, silky-tomentose;

sepals: margins villous, borders white, narrow, membranous, apex red, cuspidate, sometimes acute or obtuse, surfaces tomentose;

petals spreading, white, sometimes with hair tuft;

stamens 16–20, filaments white, anthers dark pink-purple to purple;

styles (1 or)2.

Pomes

bright to dark red, obovoid, broadly obovoid, or obconic, rarely globose or depressed-globose, 8–12.4 × 7–11.3 mm, shiny, not glaucous, sparsely pilose;

sepals flat, glabrous;

navel closed;

style remnants 3/4 from base.

red, globose, depressed-globose, subglobose, or broadly obovoid, 5–9.2 × 5.5–9.5 mm, shiny, not glaucous, villous;

sepals suberect, tomentose;

navel open;

style remnants at apex.

Pyrenes

(4 or)5.

(1 or)2.

2n

= 68 (Germany).

Cotoneaster rehderi

Cotoneaster pannosus

Phenology Flowering May–Jun; fruiting Sep–Dec. Flowering May–Aug; fruiting Oct–May.
Habitat Thickets, disturbed forests, flood plains, lakeshores Thickets, meadows, sea cliffs, canyons, coastal chaparral, springs, edges, waste ground, pastures, disturbed forests
Elevation 0–200 m (0–700 ft) 0–800 m (0–2600 ft)
Distribution
from FNA
AK; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
CA; OR; Asia (China) [Introduced in North America; introduced also in Europe, Africa (South Africa), Pacific Islands (Hawaii, New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Reports of Cotoneaster bullatus (treated as C. bullatus var. bullatus by L. Lingdi and A. R. Brach 2003) from British Columbia (J. Pojar 1999) are here referred to C. rehderi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 464. FNA vol. 9, p. 455.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. bullatus var. macrophyllus
Name authority Pojarkova: Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17: 184. (1955) Franchet: Pl. Delavay., 223. (1890)
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