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bullate cotoneaster, puckered-leaf cotoneaster

few-flower cotoneaster, Kokonor cotoneaster

Habit Shrubs, 2–5 m. Stems erect, arching; branches distichous or spiraled, maroon to brown, lenticellate, initially pilose-strigose. Shrubs, 1.5–3 m. Stems erect, becoming pendulous; branches divaricate, red-brown, initially strigose.
Leaves

deciduous;

petiole 0–3 mm, pilose-strigose;

blade elliptic, sometimes ovate-elliptic, rarely obovate, (50–)70–124(–210) x (24–)30–55(–90) mm, subcoriaceous, base cuneate, obtuse, truncate, or auriculate, margins flat, not revolute, veins 8–11, deeply sunken, apex long-acuminate, sometimes acute on short shoots, abaxial surfaces light green, initially yellowish strigose, adaxial green, shiny, not glaucous, strongly wrinkled or bulging between lateral veins (bullate), initially sparsely pilose;

fall leaves intense butter yellow and reddish purple.

deciduous;

petiole 2–3(–4) mm, pilose-strigose;

blade elliptic or obovate, sometimes broadly elliptic, broadly ovate, or suborbiculate, 8–28 × 9–23 mm, chartaceous, base rounded, margins flat, veins 3–5, superficial, apex obtuse or acute and mucronulate, abaxial surfaces pale green, initially densely pilose-strigose, adaxial initially reddish, soon bright green, dull to slightly shiny, not glaucous, flat between lateral veins, glabrous;

fall leaves turning bright red and gold.

Inflorescences

on fertile shoots 60–120 mm with 4 leaves, 8–32-flowered, lax.

on fertile shoots 15–30 mm with 3–4(–5) leaves, 1–3(or 4)-flowered, compact.

Pedicels

2–4 mm, pilose-strigose.

2–4 mm, glabrescent.

Flowers

(5–)7–9 mm, closed;

hypanthium shallowly cupulate, shiny, sparsely pilose-strigose;

sepals: margins villous, apex acute, surfaces shiny, glabrous;

petals erect-incurved, pink, red, greenish pink, or maroon, margins pink;

stamens 20–23, filaments pink, white distally, anthers white;

styles (4 or)5.

6–7 mm, open;

hypanthium widely cupulate, sparsely pilose;

sepals: margins tomentose, apex obtuse, acute, or apiculate, surfaces sparsely pilose;

petals erect-incurved, pink-tinged with purple-maroon or pink-red, base drying to purple-black, margins white, glabrous;

stamens 10–14(or 15), filaments pink, sometimes red-purple, distally dark pink, anthers pink or white, margins pale crimson, drying to dark red;

styles 2(or 3).

Pomes

bright to dark red, obovoid, broadly obovoid, or obconic, rarely globose or depressed-globose, 8–12.4 × 7–11.3 mm, shiny, not glaucous, sparsely pilose;

sepals flat, glabrous;

navel closed;

style remnants 3/4 from base.

purple-black, ellipsoid or obovoid to globose, (7–)8–9.5 × 6.5–8 mm, shiny, slightly glaucous, sparsely pilose to glabrate;

sepals erect or suberect, sparsely pilose;

navel open;

style remnants 3/4 from base.

Pyrenes

(4 or)5.

2(or 3).

2n

= 68 (Germany).

Cotoneaster rehderi

Cotoneaster nitens

Phenology Flowering May–Jun; fruiting Sep–Dec. Flowering Mar–May; fruiting Jul–Nov.
Habitat Thickets, disturbed forests, flood plains, lakeshores Thickets, edges, disturbed forests
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AK; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
OR; WA; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
Discussion

Reports of Cotoneaster bullatus (treated as C. bullatus var. bullatus by L. Lingdi and A. R. Brach 2003) from British Columbia (J. Pojar 1999) are here referred to C. rehderi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 464. FNA vol. 9, p. 463.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. bullatus var. macrophyllus
Name authority Pojarkova: Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17: 184. (1955) Rehder & E. H. Wilson: in C. S. Sargent, Pl. Wilson. 1: 156. (1912)
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