Cotoneaster rehderi |
Cotoneaster fangianus |
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bullate cotoneaster, puckered-leaf cotoneaster |
Fang's cotoneaster |
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Habit | Shrubs, 2–5 m. Stems erect, arching; branches distichous or spiraled, maroon to brown, lenticellate, initially pilose-strigose. | Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. |
Leaves | deciduous; petiole 0–3 mm, pilose-strigose; blade elliptic, sometimes ovate-elliptic, rarely obovate, (50–)70–124(–210) x (24–)30–55(–90) mm, subcoriaceous, base cuneate, obtuse, truncate, or auriculate, margins flat, not revolute, veins 8–11, deeply sunken, apex long-acuminate, sometimes acute on short shoots, abaxial surfaces light green, initially yellowish strigose, adaxial green, shiny, not glaucous, strongly wrinkled or bulging between lateral veins (bullate), initially sparsely pilose; fall leaves intense butter yellow and reddish purple. |
deciduous; petiole 2–4 mm, tomentose; blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose; fall leaves yellowish. |
Inflorescences | on fertile shoots 60–120 mm with 4 leaves, 8–32-flowered, lax. |
on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact. |
Pedicels | 2–4 mm, pilose-strigose. |
2–5 mm, tomentose-pilose. |
Flowers | (5–)7–9 mm, closed; hypanthium shallowly cupulate, shiny, sparsely pilose-strigose; sepals: margins villous, apex acute, surfaces shiny, glabrous; petals erect-incurved, pink, red, greenish pink, or maroon, margins pink; stamens 20–23, filaments pink, white distally, anthers white; styles (4 or)5. |
erect, 4–6 mm; hypanthium cupulate, densely villous; sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally; petals erect, pink or fading whitish; stamens (16–)19 or 20, filaments pale pink or white, anthers white; styles [1 or]2(or 3). |
Pomes | bright to dark red, obovoid, broadly obovoid, or obconic, rarely globose or depressed-globose, 8–12.4 × 7–11.3 mm, shiny, not glaucous, sparsely pilose; sepals flat, glabrous; navel closed; style remnants 3/4 from base. |
pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous; sepals flat, margins villous, villous; navel slightly open; style remnants 2/3 from base. |
Pyrenes | (4 or)5. |
[1 or]2(or 3). |
Cotoneaster rehderi |
Cotoneaster fangianus |
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Phenology | Flowering May–Jun; fruiting Sep–Dec. | Flowering May; fruiting Oct–Nov. |
Habitat | Thickets, disturbed forests, flood plains, lakeshores | Disturbed mesic forests |
Elevation | 0–200 m (0–700 ft) | 0–200 m (0–700 ft) |
Distribution |
AK; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe] |
OR; Asia (China) [Introduced in North America; introduced also in Europe] |
Discussion | Reports of Cotoneaster bullatus (treated as C. bullatus var. bullatus by L. Lingdi and A. R. Brach 2003) from British Columbia (J. Pojar 1999) are here referred to C. rehderi. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 464. | FNA vol. 9, p. 467. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. bullatus var. macrophyllus | |
Name authority | Pojarkova: Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17: 184. (1955) | T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) |
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