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bullate cotoneaster, puckered-leaf cotoneaster

Swedish cotoneaster

Habit Shrubs, 2–5 m. Stems erect, arching; branches distichous or spiraled, maroon to brown, lenticellate, initially pilose-strigose. Shrubs, 0.4–0.6 m. Stems ± erect or ± procumbent, arching, ends rooting; branches spiraled, green proximally, purple-black distally, initially densely strigose.
Leaves

deciduous;

petiole 0–3 mm, pilose-strigose;

blade elliptic, sometimes ovate-elliptic, rarely obovate, (50–)70–124(–210) x (24–)30–55(–90) mm, subcoriaceous, base cuneate, obtuse, truncate, or auriculate, margins flat, not revolute, veins 8–11, deeply sunken, apex long-acuminate, sometimes acute on short shoots, abaxial surfaces light green, initially yellowish strigose, adaxial green, shiny, not glaucous, strongly wrinkled or bulging between lateral veins (bullate), initially sparsely pilose;

fall leaves intense butter yellow and reddish purple.

persistent;

petiole often red, 1–3 mm, strigose;

blade elliptic, 10–27 × 4–12 mm, coriaceous, base cuneate, margins revolute, veins 4–7, superficial, apex obtuse or acute, rarely emarginate, abaxial surfaces gray-green, initially strigose-villous, midrib often red, adaxial dark green, shiny, not glaucous, flat between lateral veins, initially sparsely long-haired pilose.

Inflorescences

on fertile shoots 60–120 mm with 4 leaves, 8–32-flowered, lax.

on fertile shoots 8–40 mm with 3 or 4 leaves, 1 or 2(–6)-flowered, compact.

Pedicels

2–4 mm, pilose-strigose.

1–3.5 mm, initially sparsely pilose.

Flowers

(5–)7–9 mm, closed;

hypanthium shallowly cupulate, shiny, sparsely pilose-strigose;

sepals: margins villous, apex acute, surfaces shiny, glabrous;

petals erect-incurved, pink, red, greenish pink, or maroon, margins pink;

stamens 20–23, filaments pink, white distally, anthers white;

styles (4 or)5.

12–13 mm diam.;

buds white;

hypanthium cupulate, initially sparsely pilose;

sepals: margins villous, apex acute, surfaces villous;

petals spreading, white, glabrous;

stamens 15–20, filaments white, anthers dark red-purple;

styles 2–4(or 5).

Pomes

bright to dark red, obovoid, broadly obovoid, or obconic, rarely globose or depressed-globose, 8–12.4 × 7–11.3 mm, shiny, not glaucous, sparsely pilose;

sepals flat, glabrous;

navel closed;

style remnants 3/4 from base.

red-orange to scarlet, obovoid to subglobose, [4–]7–11 × 7–10 mm, shiny, not glaucous, glabrous or sparsely villous;

sepals depressed to slightly ascending, glabrous or sparsely villous;

navel open;

style remnants at apex.

Pyrenes

(4 or)5.

2–4(or 5) (always some with more than 2).

2n

= 34 (Germany).

Cotoneaster rehderi

Cotoneaster ×suecicus

Phenology Flowering May–Jun; fruiting Sep–Dec. Flowering May–Jun; fruiting Oct–May.
Habitat Thickets, disturbed forests, flood plains, lakeshores Hedges, edges, paths, urban waste ground
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AK; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
OR; WA; BC; QC; Europe [Introduced in North America]
[BONAP county map]
Discussion

Reports of Cotoneaster bullatus (treated as C. bullatus var. bullatus by L. Lingdi and A. R. Brach 2003) from British Columbia (J. Pojar 1999) are here referred to C. rehderi.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cotoneaster x\suecicus is of garden origin, presumably a cross between C. conspicuus and C. dammeri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 464. FNA vol. 9, p. 460.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus
Synonyms C. bullatus var. macrophyllus
Name authority Pojarkova: Bot. Mater. Gerb. Bot. Inst. Komarova Akad. Nauk S.S.S.R. 17: 184. (1955) Cotoneaster ×suecicus G. Klotz: Beitr. Phytotax. 10: 47. (1982)
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