Cotoneaster pannosus |
Rosaceae subfam. amygdaloideae |
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silver-leaf cotoneaster, woolly cotoneaster |
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Habit | Shrubs, 1–5 m. Stems erect, wide-spreading, arching, pendulous, slender; branches spiraled, purple-black, tomentose-villous. | Shrubs or trees, sometimes subshrubs or herbs. |
Leaves | persistent; petiole 4–7 mm, tomentose; blade elliptic, sometimes broadly elliptic, 15–35(–39) x 10–19(–26) mm, mid-coriaceous, base cuneate, margins flat, veins 4–6, usually superficial, sometimes slightly sunken, apex acute or obtuse, abaxial surfaces whitish tomentose, adaxial green to blue-green, dull, not glaucous, usually flat between lateral veins, initially sparsely pilose. |
alternate, sometimes opposite, simple, sometimes pinnately compound; stipules present or absent. |
Inflorescences | on fertile shoots 20–45 mm, usually with 4 leaves, (3–)5–15(–25)-flowered, compact. |
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Pedicels | 1–5 mm, tomentose. |
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Flowers | [7–]8.5–10 mm diam.; buds white; hypanthium funnelform or cupulate, silky-tomentose; sepals: margins villous, borders white, narrow, membranous, apex red, cuspidate, sometimes acute or obtuse, surfaces tomentose; petals spreading, white, sometimes with hair tuft; stamens 16–20, filaments white, anthers dark pink-purple to purple; styles (1 or)2. |
torus absent or minute; carpels 1–5(–8), distinct or +/- connate (Maleae), free or +/- adnate to hypanthium (many Maleae), styles distinct or +/- connate (some Maleae); ovules (1 or)2(–5+), collateral, clustered, or biseriate. |
Fruits | follicles aggregated or not, capsules, drupes aggregated or not, aggregated drupelets, pomes, or aggregated nutlets, rarely achenes or aggregated achenes; styles persistent or deciduous, not elongate (elongate in Gillenieae). |
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Pomes | red, globose, depressed-globose, subglobose, or broadly obovoid, 5–9.2 × 5.5–9.5 mm, shiny, not glaucous, villous; sepals suberect, tomentose; navel open; style remnants at apex. |
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Pyrenes | (1 or)2. |
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x | = 8, 9, 15, 17. |
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2n | = 68 (Germany). |
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Cotoneaster pannosus |
Rosaceae subfam. amygdaloideae |
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Phenology | Flowering May–Aug; fruiting Oct–May. | |
Habitat | Thickets, meadows, sea cliffs, canyons, coastal chaparral, springs, edges, waste ground, pastures, disturbed forests | |
Elevation | 0–800 m (0–2600 ft) | |
Distribution |
CA; OR; Asia (China) [Introduced in North America; introduced also in Europe, Africa (South Africa), Pacific Islands (Hawaii, New Zealand), Australia]
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HI; North America; Mexico; Central America; South America; Europe; Asia; Africa; Atlantic Islands (Madeira); Australia |
Discussion | Cyanogenic glycosides are usually present in Amygdaloideae; sorbitol is present. The name Amygdaloideae Arnott (1832) has priority over Spiraeoideae Arnott (1832), used by D. Potter et al. (2007), because Amygdalaceae (1820) is an earlier conserved name. Tribes 9, genera 55, species ca. 1300 (9 tribes, 38 genera, 361 species, including 20 hybrids, in the flora) (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 9, p. 455. | FNA vol. 9, p. 345. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster | Rosaceae |
Sibling taxa | ||
Subordinate taxa | ||
Name authority | Franchet: Pl. Delavay., 223. (1890) | Arnott: Botany, 107. (1832) |
Web links |