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silver-leaf cotoneaster, woolly cotoneaster

Habit Shrubs, 1–5 m. Stems erect, wide-spreading, arching, pendulous, slender; branches spiraled, purple-black, tomentose-villous. Shrubs or trees, sometimes subshrubs or herbs.
Leaves

persistent;

petiole 4–7 mm, tomentose;

blade elliptic, sometimes broadly elliptic, 15–35(–39) x 10–19(–26) mm, mid-coriaceous, base cuneate, margins flat, veins 4–6, usually superficial, sometimes slightly sunken, apex acute or obtuse, abaxial surfaces whitish tomentose, adaxial green to blue-green, dull, not glaucous, usually flat between lateral veins, initially sparsely pilose.

alternate, sometimes opposite, simple, sometimes pinnately compound;

stipules present or absent.

Inflorescences

on fertile shoots 20–45 mm, usually with 4 leaves, (3–)5–15(–25)-flowered, compact.

Pedicels

1–5 mm, tomentose.

Flowers

[7–]8.5–10 mm diam.;

buds white;

hypanthium funnelform or cupulate, silky-tomentose;

sepals: margins villous, borders white, narrow, membranous, apex red, cuspidate, sometimes acute or obtuse, surfaces tomentose;

petals spreading, white, sometimes with hair tuft;

stamens 16–20, filaments white, anthers dark pink-purple to purple;

styles (1 or)2.

torus absent or minute;

carpels 1–5(–8), distinct or +/- connate (Maleae), free or +/- adnate to hypanthium (many Maleae), styles distinct or +/- connate (some Maleae);

ovules (1 or)2(–5+), collateral, clustered, or biseriate.

Fruits

follicles aggregated or not, capsules, drupes aggregated or not, aggregated drupelets, pomes, or aggregated nutlets, rarely achenes or aggregated achenes;

styles persistent or deciduous, not elongate (elongate in Gillenieae).

Pomes

red, globose, depressed-globose, subglobose, or broadly obovoid, 5–9.2 × 5.5–9.5 mm, shiny, not glaucous, villous;

sepals suberect, tomentose;

navel open;

style remnants at apex.

Pyrenes

(1 or)2.

x

= 8, 9, 15, 17.

2n

= 68 (Germany).

Cotoneaster pannosus

Rosaceae subfam. amygdaloideae

Phenology Flowering May–Aug; fruiting Oct–May.
Habitat Thickets, meadows, sea cliffs, canyons, coastal chaparral, springs, edges, waste ground, pastures, disturbed forests
Elevation 0–800 m (0–2600 ft)
Distribution
from FNA
CA; OR; Asia (China) [Introduced in North America; introduced also in Europe, Africa (South Africa), Pacific Islands (Hawaii, New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
HI; North America; Mexico; Central America; South America; Europe; Asia; Africa; Atlantic Islands (Madeira); Australia
Discussion

Cyanogenic glycosides are usually present in Amygdaloideae; sorbitol is present.

The name Amygdaloideae Arnott (1832) has priority over Spiraeoideae Arnott (1832), used by D. Potter et al. (2007), because Amygdalaceae (1820) is an earlier conserved name.

Tribes 9, genera 55, species ca. 1300 (9 tribes, 38 genera, 361 species, including 20 hybrids, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 455. FNA vol. 9, p. 345. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Subordinate taxa
Name authority Franchet: Pl. Delavay., 223. (1890) Arnott: Botany, 107. (1832)
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