Cotoneaster pannosus |
Cotoneaster villosulus |
|
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silver-leaf cotoneaster, woolly cotoneaster |
taiping cotoneaster |
|
Habit | Shrubs, 1–5 m. Stems erect, wide-spreading, arching, pendulous, slender; branches spiraled, purple-black, tomentose-villous. | Shrubs or trees, 3–5 m. Stems erect, arching, spreading, coarse; branches distichous or spiraled, brown, initially densely villose-strigose. |
Leaves | persistent; petiole 4–7 mm, tomentose; blade elliptic, sometimes broadly elliptic, 15–35(–39) x 10–19(–26) mm, mid-coriaceous, base cuneate, margins flat, veins 4–6, usually superficial, sometimes slightly sunken, apex acute or obtuse, abaxial surfaces whitish tomentose, adaxial green to blue-green, dull, not glaucous, usually flat between lateral veins, initially sparsely pilose. |
deciduous; petiole 2–5 mm, strigose; blade elliptic or ovate, 32–72(–104) x 18–37(–46) mm, subcoriaceous to chartaceous, base obtuse or cuneate, margins flat, not revolute, veins 5–8, lightly sunken, apex attenuate to long-acuminate, often acute on short shoots, abaxial surfaces green, shiny, sparsely pilose-strigose, adaxial dark reddish green, shiny, not glaucous, soon dark green, dullish, bulging between lateral veins, rugose, pilose-strigose; fall leaves purple and rich dark red. |
Inflorescences | on fertile shoots 20–45 mm, usually with 4 leaves, (3–)5–15(–25)-flowered, compact. |
on fertile shoots 25–50 mm with 4 leaves, 3–5(–7)-flowered, mostly compact. |
Pedicels | 1–5 mm, tomentose. |
3–7 mm, densely strigose. |
Flowers | [7–]8.5–10 mm diam.; buds white; hypanthium funnelform or cupulate, silky-tomentose; sepals: margins villous, borders white, narrow, membranous, apex red, cuspidate, sometimes acute or obtuse, surfaces tomentose; petals spreading, white, sometimes with hair tuft; stamens 16–20, filaments white, anthers dark pink-purple to purple; styles (1 or)2. |
6–8 mm, opening small; hypanthium cupulate, tomentose-strigose; sepals: margins tomentose, borders brown, broad, glabrous, apex acute, obtuse, or apiculate, surfaces strigose; petals erect-incurved, pink or red, base dark red, margins white; stamens 18–20, filaments pale pink, white distally, anthers white; styles 2 or 3. |
Pomes | red, globose, depressed-globose, subglobose, or broadly obovoid, 5–9.2 × 5.5–9.5 mm, shiny, not glaucous, villous; sepals suberect, tomentose; navel open; style remnants at apex. |
purple-black, broadly ellipsoid to broadly obovoid, rarely cylindric, 9–11 × 8–11 mm, slightly shiny, not or slightly glaucous, villous; sepals flat or ascending, margins tomentose, strigose; forming star over open navel; style remnants 4/5 from base. |
Pyrenes | (1 or)2. |
2 or 3. |
2n | = 68 (Germany). |
= 68 (Germany). |
Cotoneaster pannosus |
Cotoneaster villosulus |
|
Phenology | Flowering May–Aug; fruiting Oct–May. | Flowering May (often reflowering Sep); fruiting Aug–Nov. |
Habitat | Thickets, meadows, sea cliffs, canyons, coastal chaparral, springs, edges, waste ground, pastures, disturbed forests | Thickets, open forests |
Elevation | 0–800 m (0–2600 ft) | 0–50 m (0–200 ft) |
Distribution |
CA; OR; Asia (China) [Introduced in North America; introduced also in Europe, Africa (South Africa), Pacific Islands (Hawaii, New Zealand), Australia]
|
WA; Asia (China) [Introduced in North America; introduced also in Europe] |
Discussion | The difficulties of typification and taxonomy for black-fruited species were discussed by J. Fryer and B. Hylmö (1997). Some collections of Cotoneaster villosulus from eastern Washington need study. They approach the very similar C. hsingshangensis J. Fryer & B. Hylmö, a Chinese species with larger, more wrinkled leaves, more globose pomes, and less densely hairy flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 9, p. 455. | FNA vol. 9, p. 464. |
Parent taxa | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster | Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster |
Sibling taxa | ||
Synonyms | C. acutifolius var. villosulus | |
Name authority | Franchet: Pl. Delavay., 223. (1890) | (Rehder & E. H. Wilson) Flinck & B. Hylmö: Bot. Not. 115: 383. (1962) |
Web links |