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silver-leaf cotoneaster, woolly cotoneaster

taiping cotoneaster

Habit Shrubs, 1–5 m. Stems erect, wide-spreading, arching, pendulous, slender; branches spiraled, purple-black, tomentose-villous. Shrubs or trees, 3–5 m. Stems erect, arching, spreading, coarse; branches distichous or spiraled, brown, initially densely villose-strigose.
Leaves

persistent;

petiole 4–7 mm, tomentose;

blade elliptic, sometimes broadly elliptic, 15–35(–39) x 10–19(–26) mm, mid-coriaceous, base cuneate, margins flat, veins 4–6, usually superficial, sometimes slightly sunken, apex acute or obtuse, abaxial surfaces whitish tomentose, adaxial green to blue-green, dull, not glaucous, usually flat between lateral veins, initially sparsely pilose.

deciduous;

petiole 2–5 mm, strigose;

blade elliptic or ovate, 32–72(–104) x 18–37(–46) mm, subcoriaceous to chartaceous, base obtuse or cuneate, margins flat, not revolute, veins 5–8, lightly sunken, apex attenuate to long-acuminate, often acute on short shoots, abaxial surfaces green, shiny, sparsely pilose-strigose, adaxial dark reddish green, shiny, not glaucous, soon dark green, dullish, bulging between lateral veins, rugose, pilose-strigose;

fall leaves purple and rich dark red.

Inflorescences

on fertile shoots 20–45 mm, usually with 4 leaves, (3–)5–15(–25)-flowered, compact.

on fertile shoots 25–50 mm with 4 leaves, 3–5(–7)-flowered, mostly compact.

Pedicels

1–5 mm, tomentose.

3–7 mm, densely strigose.

Flowers

[7–]8.5–10 mm diam.;

buds white;

hypanthium funnelform or cupulate, silky-tomentose;

sepals: margins villous, borders white, narrow, membranous, apex red, cuspidate, sometimes acute or obtuse, surfaces tomentose;

petals spreading, white, sometimes with hair tuft;

stamens 16–20, filaments white, anthers dark pink-purple to purple;

styles (1 or)2.

6–8 mm, opening small;

hypanthium cupulate, tomentose-strigose;

sepals: margins tomentose, borders brown, broad, glabrous, apex acute, obtuse, or apiculate, surfaces strigose;

petals erect-incurved, pink or red, base dark red, margins white;

stamens 18–20, filaments pale pink, white distally, anthers white;

styles 2 or 3.

Pomes

red, globose, depressed-globose, subglobose, or broadly obovoid, 5–9.2 × 5.5–9.5 mm, shiny, not glaucous, villous;

sepals suberect, tomentose;

navel open;

style remnants at apex.

purple-black, broadly ellipsoid to broadly obovoid, rarely cylindric, 9–11 × 8–11 mm, slightly shiny, not or slightly glaucous, villous;

sepals flat or ascending, margins tomentose, strigose; forming star over open navel;

style remnants 4/5 from base.

Pyrenes

(1 or)2.

2 or 3.

2n

= 68 (Germany).

= 68 (Germany).

Cotoneaster pannosus

Cotoneaster villosulus

Phenology Flowering May–Aug; fruiting Oct–May. Flowering May (often reflowering Sep); fruiting Aug–Nov.
Habitat Thickets, meadows, sea cliffs, canyons, coastal chaparral, springs, edges, waste ground, pastures, disturbed forests Thickets, open forests
Elevation 0–800 m (0–2600 ft) 0–50 m (0–200 ft)
Distribution
from FNA
CA; OR; Asia (China) [Introduced in North America; introduced also in Europe, Africa (South Africa), Pacific Islands (Hawaii, New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
Discussion

The difficulties of typification and taxonomy for black-fruited species were discussed by J. Fryer and B. Hylmö (1997). Some collections of Cotoneaster villosulus from eastern Washington need study. They approach the very similar C. hsingshangensis J. Fryer & B. Hylmö, a Chinese species with larger, more wrinkled leaves, more globose pomes, and less densely hairy flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 455. FNA vol. 9, p. 464.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. ×suecicus
Synonyms C. acutifolius var. villosulus
Name authority Franchet: Pl. Delavay., 223. (1890) (Rehder & E. H. Wilson) Flinck & B. Hylmö: Bot. Not. 115: 383. (1962)
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