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one-stone cotoneaster

Habit Shrubs or trees, 3–5 m. Stems loosely erect, arching, spreading; branches spiraled, maroon, lenticellate, initially pilose-strigose. Shrubs or trees, sometimes subshrubs or herbs.
Leaves

deciduous;

petiole 5–9 mm, pilose-strigose;

blade elliptic, broadly elliptic, or broadly obovate, sometimes suborbiculate, 25–58 × 15–40 mm, subcoriaceous, base obtuse or cuneate, margins flat, veins 4–6, slightly sunken, apex obtuse or truncate, seldom acute, abaxial surfaces color not recorded, reticulate, tomentose-pilose, later thinning, adaxial dark green, dull, coating not recorded, flat between lateral veins, glabrescent;

fall leaves pale yellowish green.

alternate, sometimes opposite, simple, sometimes pinnately compound;

stipules present or absent.

Inflorescences

on fertile shoots 30–50 mm with 3–4 leaves, 7–20-flowered, lax.

Pedicels

2–5 mm, pilose-strigose.

Flowers

10–12 mm diam., buds white, hypanthium campanulate to cupulate, sparsely pilose-strigose;

sepals: margins reddish, villous, borders maroon-tipped, membranous, apex acute, acuminate, or obtuse, surfaces sparsely pilose-strigose;

petals spreading, white, sometimes with hair tuft;

stamens (15–)20, filaments white;

anthers purple to blackish purple;

styles 1(or 2).

torus absent or minute;

carpels 1–5(–8), distinct or +/- connate (Maleae), free or +/- adnate to hypanthium (many Maleae), styles distinct or +/- connate (some Maleae);

ovules (1 or)2(–5+), collateral, clustered, or biseriate.

Fruits

follicles aggregated or not, capsules, drupes aggregated or not, aggregated drupelets, pomes, or aggregated nutlets, rarely achenes or aggregated achenes;

styles persistent or deciduous, not elongate (elongate in Gillenieae).

Pomes

dark red or ruby to maroon, maturing dark purple or purple-black, obovoid, rarely globose or ellipsoid, 9–12.5 × 8.5–11.5 mm, dull, glaucous, glabrous;

sepals flat, margins sparsely hairy, sparsely hairy to glabrate;

navel open;

style remnants at apex.

Pyrenes

1(or 2).

x

= 8, 9, 15, 17.

Cotoneaster monopyrenus

Rosaceae subfam. amygdaloideae

Phenology Flowering May–Jun; fruiting Jul–Nov.
Habitat Thickets
Elevation 0–200 m (0–700 ft)
Distribution
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
from FNA
HI; North America; Mexico; Central America; South America; Europe; Asia; Africa; Atlantic Islands (Madeira); Australia
Discussion

L. Lingdi and A. R. Brach (2003) synonymized Cotoneaster monopyrenus with C. hebephyllus (as C. hebephyllus var. hebephyllus).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cyanogenic glycosides are usually present in Amygdaloideae; sorbitol is present.

The name Amygdaloideae Arnott (1832) has priority over Spiraeoideae Arnott (1832), used by D. Potter et al. (2007), because Amygdalaceae (1820) is an earlier conserved name.

Tribes 9, genera 55, species ca. 1300 (9 tribes, 38 genera, 361 species, including 20 hybrids, in the flora)

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 457. FNA vol. 9, p. 345. Author: Luc Brouillet.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Subordinate taxa
Synonyms C. hebephyllus var. monopyrenus
Name authority (W. W. Smith) Flinck & B. Hylmö: Bot. Not. 119: 459. (1966) Arnott: Botany, 107. (1832)
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