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one-stone cotoneaster

Fang's cotoneaster

Habit Shrubs or trees, 3–5 m. Stems loosely erect, arching, spreading; branches spiraled, maroon, lenticellate, initially pilose-strigose. Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose.
Leaves

deciduous;

petiole 5–9 mm, pilose-strigose;

blade elliptic, broadly elliptic, or broadly obovate, sometimes suborbiculate, 25–58 × 15–40 mm, subcoriaceous, base obtuse or cuneate, margins flat, veins 4–6, slightly sunken, apex obtuse or truncate, seldom acute, abaxial surfaces color not recorded, reticulate, tomentose-pilose, later thinning, adaxial dark green, dull, coating not recorded, flat between lateral veins, glabrescent;

fall leaves pale yellowish green.

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

Inflorescences

on fertile shoots 30–50 mm with 3–4 leaves, 7–20-flowered, lax.

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

Pedicels

2–5 mm, pilose-strigose.

2–5 mm, tomentose-pilose.

Flowers

10–12 mm diam., buds white, hypanthium campanulate to cupulate, sparsely pilose-strigose;

sepals: margins reddish, villous, borders maroon-tipped, membranous, apex acute, acuminate, or obtuse, surfaces sparsely pilose-strigose;

petals spreading, white, sometimes with hair tuft;

stamens (15–)20, filaments white;

anthers purple to blackish purple;

styles 1(or 2).

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

Pomes

dark red or ruby to maroon, maturing dark purple or purple-black, obovoid, rarely globose or ellipsoid, 9–12.5 × 8.5–11.5 mm, dull, glaucous, glabrous;

sepals flat, margins sparsely hairy, sparsely hairy to glabrate;

navel open;

style remnants at apex.

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

Pyrenes

1(or 2).

[1 or]2(or 3).

Cotoneaster monopyrenus

Cotoneaster fangianus

Phenology Flowering May–Jun; fruiting Jul–Nov. Flowering May; fruiting Oct–Nov.
Habitat Thickets Disturbed mesic forests
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
Discussion

L. Lingdi and A. R. Brach (2003) synonymized Cotoneaster monopyrenus with C. hebephyllus (as C. hebephyllus var. hebephyllus).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 457. FNA vol. 9, p. 467.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. hebephyllus var. monopyrenus
Name authority (W. W. Smith) Flinck & B. Hylmö: Bot. Not. 119: 459. (1966) T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963)
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