Cotoneaster monopyrenus |
Cotoneaster divaricatus |
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one-stone cotoneaster |
bearberry cotoneaster, spreading cotoneaster, stretch-fruit cotoneaster |
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Habit | Shrubs or trees, 3–5 m. Stems loosely erect, arching, spreading; branches spiraled, maroon, lenticellate, initially pilose-strigose. | Shrubs, 1.5–2 m; crown spreading, rounded. |
Stems | erect; branches divaricate or spiraled, spreading, lateral branches well developed, usually arching, long, brownish violet, initially strigose. |
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Leaves | deciduous; petiole 5–9 mm, pilose-strigose; blade elliptic, broadly elliptic, or broadly obovate, sometimes suborbiculate, 25–58 × 15–40 mm, subcoriaceous, base obtuse or cuneate, margins flat, veins 4–6, slightly sunken, apex obtuse or truncate, seldom acute, abaxial surfaces color not recorded, reticulate, tomentose-pilose, later thinning, adaxial dark green, dull, coating not recorded, flat between lateral veins, glabrescent; fall leaves pale yellowish green. |
deciduous; petiole 3–4 mm, strigose; blade elliptic, rarely ovate or suborbiculate, on sterile shoots usually elliptic or broadly elliptic, 10–30 × 7–21 mm, chartaceous, base obtuse, margins flat, not revolute, sometimes undulate, veins 3 or 4, superficial or slightly sunken, apex acute, rarely acuminate, abaxial surfaces pale green, initially strigose, adaxial dark green, shiny, not glaucous, flat between lateral veins, initially sparsely strigose; fall leaves orange, red, or reddish purple. |
Inflorescences | on fertile shoots 30–50 mm with 3–4 leaves, 7–20-flowered, lax. |
on fertile shoots 10–20 mm with 4 leaves, 1–4-flowered, compact. |
Pedicels | 2–5 mm, pilose-strigose. |
1–3 mm, strigose. |
Flowers | 10–12 mm diam., buds white, hypanthium campanulate to cupulate, sparsely pilose-strigose; sepals: margins reddish, villous, borders maroon-tipped, membranous, apex acute, acuminate, or obtuse, surfaces sparsely pilose-strigose; petals spreading, white, sometimes with hair tuft; stamens (15–)20, filaments white; anthers purple to blackish purple; styles 1(or 2). |
6–8 mm, opening small; hypanthium cupulate, strigose; sepals: margins villous, apex acute or acuminate, surfaces strigose; petals erect-incurved, pale red, base dark red, border white; stamens 10–15, filaments dark red, whitish distally, anthers white; styles (1 or)2(or 3). |
Pomes | dark red or ruby to maroon, maturing dark purple or purple-black, obovoid, rarely globose or ellipsoid, 9–12.5 × 8.5–11.5 mm, dull, glaucous, glabrous; sepals flat, margins sparsely hairy, sparsely hairy to glabrate; navel open; style remnants at apex. |
dark red to ruby (finally blackish red), cylindric, oblong-ellipsoid, or narrowly obovoid, 8–11 × 4–8 mm, shiny, not glaucous, sparsely pilose; sepals suberect, sparsely pilose; navel open; style remnants 2/3 from base. |
Pyrenes | 1(or 2). |
(1 or)2(or 3) [rarely 4]. |
Cotoneaster monopyrenus |
Cotoneaster divaricatus |
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Phenology | Flowering May–Jun; fruiting Jul–Nov. | Flowering Apr–May; fruiting Sep–Dec. |
Habitat | Thickets | Thickets, edges, disturbed forests, flood plains, rarely epiphytic on maple |
Elevation | 0–200 m (0–700 ft) | 0–700 m (0–2300 ft) |
Distribution |
WA; Asia (China) [Introduced in North America; introduced also in Europe] |
IL; IN; KY; MA; MI; NY; OH; OR; PA; WA; WI; BC; ON; Asia (China) [Introduced in North America; introduced also in Europe]
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Discussion | L. Lingdi and A. R. Brach (2003) synonymized Cotoneaster monopyrenus with C. hebephyllus (as C. hebephyllus var. hebephyllus). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although rarely collected, Cotoneaster divaricatus is the most widely escaped cotoneaster in eastern North America. Misidentified specimens of it have been reported from New York as C. hupehensis Rehder & E. H. Wilson (R. S. Mitchell and G. C. Tucker 1997), from Illinois as C. acutifolius (J. T. Kartesz and C. A. Meacham 2003), from Indiana and Ontario as C. horizontalis (Kartesz and Meacham), from Kentucky as C. apiculatus Rehder & E. H. Wilson (Kartesz and Meacham), and from Wisconsin as C. lucidus (Kartesz and Meacham). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 457. | FNA vol. 9, p. 461. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. hebephyllus var. monopyrenus | |
Name authority | (W. W. Smith) Flinck & B. Hylmö: Bot. Not. 119: 459. (1966) | Rehder & E. H. Wilson: in C. S. Sargent, Pl. Wilson. 1: 157. (1912) |
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