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dark cotoneaster, dark-fruit cotoneaster

cotoneaster, Tengyueh cotoneaster

Habit Shrubs, 2–2.5 m. Stems loosely erect; branches spiraled, yellow brown to red brown, lenticellate, shiny, initially densely pilose. Shrubs, 2–2.5 m; crown ± rounded.
Stems

erect, arching, long;

branches distichous, grayish maroon, initially densely strigose.

Leaves

deciduous;

petiole 4–7 mm, tomentose-villous;

blade elliptic to ovate, 33–45 × 20–32 mm, chartaceous, base rounded, margins flat, veins 5–7, superficial, apex acute or obtuse, abaxial surfaces densely silvery-pilose-villous, adaxial green to dark green, dull to slightly shiny, not glaucous, flat between lateral veins, rugose, sparsely pilose;

fall leaves lacking notable color.

deciduous (sometimes tardily so on vigorous shoots);

petiole 1–4 mm, strigose;

blade ovate or elliptic, (25–)30–50(–61) x 12–32 mm, chartaceous, base cuneate or rounded, margins flat, veins 3–5, deeply sunken, apex acuminate or acute, abaxial surfaces grayish green, villose-strigose or sparsely to moderately gray-tomentose, adaxial green to dark green, shiny, not glaucous, flat or faintly bulging between lateral veins, pilose-strigose;

fall leaves yellow to orange.

Inflorescences

on fertile shoots 25–30 mm with 3 or 4 leaves, 5–13-flowered, pendent, lax.

on fertile shoots 25–40 mm with (3 or)4 leaves, 3–7(–9)-flowered, compact.

Pedicels

3–8 mm, sometimes sparsely villous.

1–4 mm, densely strigose.

Flowers

erect, 6–7 mm, open;

hypanthium cupulate, dark reddish brown, glabrous;

sepals: margins erose, apex acute or obtuse, surfaces glabrous, often apically villous;

petals erect, greenish white with pink, red, base slightly darkened, margins white, glabrous;

stamens 20(–22), filaments white, anthers white;

styles 2–4.

erect or ascending, [5–]6–8 mm, opening small;

hypanthium cupulate, strigose;

sepals: margins villous, borders reddish brown, glabrous, apex acuminate or cuspidate, surfaces strigose;

petals erect-incurved, pink to red, base dark pink or red, rarely dark red or maroon, margins white;

stamens 20 or 21, filaments pink or pale pink, whitish distally, anthers white;

styles (2 or)3–5.

Pomes

purple-black, obovoid or globose, 7–9 × 7–9 mm, dull, glaucous with blue tinge, glabrous;

sepals suberect, glabrous or apically villous;

navel open;

style remnants 2/3 from base.

spreading or pendent, bright red, broadly obovoid or subglobose, rarely oblong-obovoid, 6.3–10.4 × 5.8–9.1 mm, shiny, not glaucous, sparsely strigose;

sepals flat or suberect, strigose;

navel closed;

style remnants 2/3 from base.

Pyrenes

2–4.

(2 or)3–5.

2n

= 48–52, 68 (Russia).

= 68 (Germany).

Cotoneaster melanocarpus

Cotoneaster tengyuehensis

Phenology Flowering Apr–May; fruiting Jul–Aug. Flowering May–Jun; fruiting Sep–Feb.
Habitat Forested ravines Thickets, creek banks, open forests, edges
Elevation 300–500 m (1000–1600 ft) 0–50 m (0–200 ft)
Distribution
from FNA
MB; Eurasia (Russia, Ukraine) [Introduced in North America]
[BONAP county map]
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
Discussion

Cotoneaster melanocarpus is said to hybridize spontaneously with Sorbus aucuparia in Siberia, forming x\Sorbocotoneaster Pojarkova. The actual Cotoneaster parent is more likely to be a diploid and needs investigation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. Lingdi and A. R. Brach (2003) confused Cotoneaster tengyuehensis with C. glomerulatus W. W. Smith, a species with small fruits 5 mm in diameter, with 5 pyrenes and styles, and slightly smaller leaves to 40–50 mm. In the flora area, C. tengyuehensis is more likely to be mistaken for C. franchetii, which is a true evergreen with orange-red, obovate to obconic fruits, and pink to purple stamens. Naturalized shrubs of C. tengyuehensis drop their foliage in midwinter.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 465.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. vulgaris var. melanocarpus
Name authority (Ledebour) Loddiges: Fam. Nat. Syn. Monogr. 3: 223. (1847) J. Fryer & B. Hylmö: Watsonia 21: 338. (1997)
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