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dark cotoneaster, dark-fruit cotoneaster

cotoneaster, crisp's cotoneaster

Habit Shrubs, 2–2.5 m. Stems loosely erect; branches spiraled, yellow brown to red brown, lenticellate, shiny, initially densely pilose. Shrubs or trees, 1.5–6 m. Stems narrowly erect, arching; branches spiraled, purple-black, initially silky tomentose.
Leaves

deciduous;

petiole 4–7 mm, tomentose-villous;

blade elliptic to ovate, 33–45 × 20–32 mm, chartaceous, base rounded, margins flat, veins 5–7, superficial, apex acute or obtuse, abaxial surfaces densely silvery-pilose-villous, adaxial green to dark green, dull to slightly shiny, not glaucous, flat between lateral veins, rugose, sparsely pilose;

fall leaves lacking notable color.

persistent, rarely semipersistent (in harsh winters);

petiole 5–7 mm, tomentose;

blade elliptic, sometimes ovate, 25–46 × 11–28 mm, subcoriaceous or coriaceous, base cuneate, margins flat, veins 4–7, superficial, rarely faintly sunken, apex acute, abaxial surfaces initially whitish tomentose, becoming glabrate, adaxial green to dark green, dull, slightly glaucous, flat between lateral veins, initially sparsely pilose.

Inflorescences

on fertile shoots 25–30 mm with 3 or 4 leaves, 5–13-flowered, pendent, lax.

on fertile shoots 40–70 mm with 4 leaves, 7–25-flowered, compact to ± lax.

Pedicels

3–8 mm, sometimes sparsely villous.

1–7 mm, silky tomentose.

Flowers

erect, 6–7 mm, open;

hypanthium cupulate, dark reddish brown, glabrous;

sepals: margins erose, apex acute or obtuse, surfaces glabrous, often apically villous;

petals erect, greenish white with pink, red, base slightly darkened, margins white, glabrous;

stamens 20(–22), filaments white, anthers white;

styles 2–4.

(7–)10–11 mm diam.;

buds white;

hypanthium funnelform, silky tomentose;

sepals: margins villous, apex cuspidate or acuminate, surfaces silky tomentose;

petals spreading, white, rarely with hair tuft;

stamens 20, filaments white, anthers reddish purple;

styles (1 or)2.

Pomes

purple-black, obovoid or globose, 7–9 × 7–9 mm, dull, glaucous with blue tinge, glabrous;

sepals suberect, glabrous or apically villous;

navel open;

style remnants 2/3 from base.

red to dark red, globose or subglobose, 6–8 × 6–8 mm, dull or slightly shiny, slightly glaucous, glabrescent;

sepals flat, depressed, or incurved, densely villous;

navel open;

style remnants at apex.

Pyrenes

2–4.

(1 or)2.

2n

= 48–52, 68 (Russia).

= 51 (Germany).

Cotoneaster melanocarpus

Cotoneaster crispii

Phenology Flowering Apr–May; fruiting Jul–Aug. Flowering Jun–Jul; fruiting Oct–Feb.
Habitat Forested ravines Thickets, building edges, cracks in pavement
Elevation 300–500 m (1000–1600 ft) 0–200 m (0–700 ft)
Distribution
from FNA
MB; Eurasia (Russia, Ukraine) [Introduced in North America]
[BONAP county map]
from FNA
BC; Asia (China) [Introduced in North America]
[BONAP county map]
Discussion

Cotoneaster melanocarpus is said to hybridize spontaneously with Sorbus aucuparia in Siberia, forming x\Sorbocotoneaster Pojarkova. The actual Cotoneaster parent is more likely to be a diploid and needs investigation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cotoneaster crispii was collected recently wild in Yunnan. The species is often treated as a hybrid of garden origin between C. frigidus and C. pannosus. Plants of it differ from those of C. frigidus in their smaller and persistent leaves, and it has larger leaves and fruits than C. pannosus. The species was overlooked in the Flora of China (L. Lingdi and A. R. Brach 2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 454.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. vulgaris var. melanocarpus
Name authority (Ledebour) Loddiges: Fam. Nat. Syn. Monogr. 3: 223. (1847) Exell: Gard. Chron., ser. 3, 83: 44. (1928) — as hybrid
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