Cotoneaster lucidus |
Cotoneaster villosulus |
|
---|---|---|
cotonéaster à feuilles aiguës, hedge cotoneaster, shiny cotoneaster |
taiping cotoneaster |
|
Habit | Shrubs, 1.5–2.5 m. Stems erect, arching, or spreading; branches spiraled or distichous, dense, greenish gray to maroon, not lenticellate, initially strigose. | Shrubs or trees, 3–5 m. Stems erect, arching, spreading, coarse; branches distichous or spiraled, brown, initially densely villose-strigose. |
Leaves | deciduous; petiole 4–8 mm, strigose; blade elliptic or ovate, sometimes obovate, 28–50(–65) x 17–32(–40) mm, chartaceous, base cuneate or rounded, margins flat, not revolute, veins 3–5, sunken, apex short-acuminate or acute, abaxial surfaces green, sparsely pilose-strigose, adaxial dark green, intensely shiny, not glaucous, flat or scarcely bulging between lateral veins, slightly rugose, initially sparsely strigose; fall leaves intense dark red, yellow, or orange. |
deciduous; petiole 2–5 mm, strigose; blade elliptic or ovate, 32–72(–104) x 18–37(–46) mm, subcoriaceous to chartaceous, base obtuse or cuneate, margins flat, not revolute, veins 5–8, lightly sunken, apex attenuate to long-acuminate, often acute on short shoots, abaxial surfaces green, shiny, sparsely pilose-strigose, adaxial dark reddish green, shiny, not glaucous, soon dark green, dullish, bulging between lateral veins, rugose, pilose-strigose; fall leaves purple and rich dark red. |
Inflorescences | on fertile shoots 25–50 mm with 4 or 5 leaves, 2–6(–15)-flowered, lax. |
on fertile shoots 25–50 mm with 4 leaves, 3–5(–7)-flowered, mostly compact. |
Pedicels | 5–12 mm, sparsely pilose. |
3–7 mm, densely strigose. |
Flowers | 5.5–8 mm, largely open; hypanthium cupulate, base sparsely pilose, glabrescent; sepals: margins villous, apex acute or acuminate, surfaces glabrous; petals erect, pinkish white, sometimes green-tinged, base dark pink or reddish; stamens 18–20, filaments pink, base dark pink, anthers white; styles 2 or 3. |
6–8 mm, opening small; hypanthium cupulate, tomentose-strigose; sepals: margins tomentose, borders brown, broad, glabrous, apex acute, obtuse, or apiculate, surfaces strigose; petals erect-incurved, pink or red, base dark red, margins white; stamens 18–20, filaments pale pink, white distally, anthers white; styles 2 or 3. |
Pomes | black, broadly obovoid to obovoid, rarely ellipsoid or globose, (8–)9.5–12.1 x (7.5–)8.7–11.8 mm, shiny, glaucous, sparsely pilose; sepals flat, margins villous, sparsely pilose; navel open; style remnants 2/3 from base. |
purple-black, broadly ellipsoid to broadly obovoid, rarely cylindric, 9–11 × 8–11 mm, slightly shiny, not or slightly glaucous, villous; sepals flat or ascending, margins tomentose, strigose; forming star over open navel; style remnants 4/5 from base. |
Pyrenes | 2 or 3. |
2 or 3. |
2n | = 68 (Germany). |
= 68 (Germany). |
Cotoneaster lucidus |
Cotoneaster villosulus |
|
Phenology | Flowering Mar–Jul (sometimes reflowering in Sep); fruiting Jul–Oct. | Flowering May (often reflowering Sep); fruiting Aug–Nov. |
Habitat | Deciduous or coniferous forests, ravines, lakeshores, forested dunes, thickets, edges | Thickets, open forests |
Elevation | 0–2300 m (0–7500 ft) | 0–50 m (0–200 ft) |
Distribution |
CO; ID; IN; MN; WA; WY; AB; BC; MB; ON; QC; SK; Asia (Mongolia, Siberian Russia) [Introduced in North America; introduced also in Europe] |
WA; Asia (China) [Introduced in North America; introduced also in Europe] |
Discussion | Leaf glossiness is often retained on herbarium specimens and is visible with strong lighting and magnification. Plants of Cotoneaster acutifolius of China and Mongolia are seldom seen in cultivation. Reports of it naturalized in North America are here referred to the commonly cultivated C. lucidus, which tolerates continental winters. No specimens have been seen to support reports of the latter from Montana and Oregon. Some reports of C. melanocarpus from Alberta and Manitoba (H. J. Scoggan 1978–1979, part 3) are here referred to C. lucidus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The difficulties of typification and taxonomy for black-fruited species were discussed by J. Fryer and B. Hylmö (1997). Some collections of Cotoneaster villosulus from eastern Washington need study. They approach the very similar C. hsingshangensis J. Fryer & B. Hylmö, a Chinese species with larger, more wrinkled leaves, more globose pomes, and less densely hairy flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 463. | FNA vol. 9, p. 464. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. acutifolius var. lucidus | C. acutifolius var. villosulus |
Name authority | Schlechtendal: Linnaea 27: 541. (1854) | (Rehder & E. H. Wilson) Flinck & B. Hylmö: Bot. Not. 115: 383. (1962) |
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