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cotoneaster, Franchet's cotoneaster, orange cotoneaster

entire-leaf cotoneaster, small-leaf cotoneaster

Habit Shrubs, 2–3 m. Stems erect, arching, lax, slender; branches distichous, maroon, initially densely pilose-strigose. Shrubs, wide-spreading, 0.5–1.5 m. Stems weakly ascending, suberect, or decumbent; branches spiraled, purple-black, sometimes green and brown striate, initially strigose.
Leaves

persistent, often 30–40 mm apart;

petiole 2–4 mm, tomentose-pilose;

blade ovate or elliptic, 22–58 × 12–29 mm, coriaceous, base cuneate or obtuse, margins flat, veins 4 or 5, sunken, apex acute or acuminate, mucronate, abaxial surfaces silvery-tomentose, adaxial grayish green, slightly shiny, not glaucous, flat, rarely slightly bulging between lateral veins, drying slightly rugose, pilose.

persistent, petiole 1–5 mm, strigose;

blade oblanceolate or oblong, rarely lanceolate, 8–17 × 3–8 mm, coriaceous, base cuneate, margins revolute, veins 2–4, superficial, apex rotund or obtuse, rarely emarginate or acute, abaxial surfaces grayish white, faintly reticulate, initially densely strigose or strigose-pilose, later glabrate, adaxial dark green, sometimes blue-green, shiny, not glaucous, not bulging between lateral veins, initially strigose, soon glabrate.

Inflorescences

on fertile shoots 30–60 mm with 4 leaves, 5–15(–25)-flowered, compact.

on fertile shoots 5–12 mm with 2 or 3 leaves, 1(or 2)-flowered, subsessile.

Pedicels

1–4 mm, densely long-silky hairy.

erect, densely strigose.

Flowers

(5–)7–8(–10) mm, opening small;

hypanthium cupulate, densely long-silky hairy;

sepals: margins villous, borders purple, glabrous, apex cuspidate or acuminate, surfaces silky hairy;

petals erect-incurved, pink or red, base dark pink to dark red, margins off-white;

stamens 16–21, filaments pink, distally white, anthers pink or purplish pink;

styles 2 or 3(or 4).

7–15 mm diam.;

buds pink;

hypanthium narrowly cupulate, densely strigose;

sepals: margins densely villose-strigose, borders membranous, apex often red, acute or obtuse, sometimes shortly mucronate, surfaces often densely strigose-villous;

petals spreading, white, glabrous;

stamens 20, filaments white, anthers red-purple;

styles 2[or 3].

Pomes

orange-red, obovoid base rounded or obconic base cuneate, straight-sided, 8–12 × 6–9 mm, apex flat, shiny, not glaucous, sparsely pilose;

sepals flat, often asymmetric with 2 sepals longer than others, tomentose;

navel slightly open;

style remnants 3/4 from base.

dark red with crimson, sometimes red, depressed-globose, 7–9 × 7–10 mm, dull or slightly shiny, glaucous, sparsely villous;

sepals suberect or erect, villous;

navel open;

style remnants at apex.

Pyrenes

2 or 3(or 4).

2[or 3].

2n

= 68 (Germany).

= 68 (Germany).

Cotoneaster franchetii

Cotoneaster integrifolius

Phenology Flowering May–Jul; fruiting Oct–Apr. Flowering Apr–Jun; fruiting Aug–Mar.
Habitat Edges, thickets, disturbed forests, flood plains, urban waste ground, mossy ledges, wet prairie remnants Openings in conifer forests
Elevation 0–700 m (0–2300 ft) 0–200 m (0–700 ft)
Distribution
from FNA
CA; OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe, Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Asia [Introduced in North America; introduced also in Europe]
Discussion

Reports of Cotoneaster franchetii from Colorado (J. Ells 2006) are here referred to C. lucidus. A few specimens escaped in Washington have red filaments, not pink, but otherwise agree with C. franchetii. Other escapes in Washington suggest the closely related C. wardii W. W. Smith, distinguished by semievergreen habit, longer fertile shoots, more lax and elongate inflorescences, and white anthers. The entire complex needs biosystematic review.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cotoneaster integrifolius was treated as C. microphyllus Wallich ex Lindley var. thymifolius (Baker) Koehne by L. Lingdi and A. R. Brach (2003).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 465. FNA vol. 9, p. 458.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms Crataegus integrifolius
Name authority Bois: Rev. Hort. 74: 380, figs. 159–161, 164. (1902) (Roxburgh) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 12: 779. (1963)
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