Cotoneaster fangianus |
Cotoneaster villosulus |
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Fang's cotoneaster |
taiping cotoneaster |
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Habit | Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. | Shrubs or trees, 3–5 m. Stems erect, arching, spreading, coarse; branches distichous or spiraled, brown, initially densely villose-strigose. |
Leaves | deciduous; petiole 2–4 mm, tomentose; blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose; fall leaves yellowish. |
deciduous; petiole 2–5 mm, strigose; blade elliptic or ovate, 32–72(–104) x 18–37(–46) mm, subcoriaceous to chartaceous, base obtuse or cuneate, margins flat, not revolute, veins 5–8, lightly sunken, apex attenuate to long-acuminate, often acute on short shoots, abaxial surfaces green, shiny, sparsely pilose-strigose, adaxial dark reddish green, shiny, not glaucous, soon dark green, dullish, bulging between lateral veins, rugose, pilose-strigose; fall leaves purple and rich dark red. |
Inflorescences | on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact. |
on fertile shoots 25–50 mm with 4 leaves, 3–5(–7)-flowered, mostly compact. |
Pedicels | 2–5 mm, tomentose-pilose. |
3–7 mm, densely strigose. |
Flowers | erect, 4–6 mm; hypanthium cupulate, densely villous; sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally; petals erect, pink or fading whitish; stamens (16–)19 or 20, filaments pale pink or white, anthers white; styles [1 or]2(or 3). |
6–8 mm, opening small; hypanthium cupulate, tomentose-strigose; sepals: margins tomentose, borders brown, broad, glabrous, apex acute, obtuse, or apiculate, surfaces strigose; petals erect-incurved, pink or red, base dark red, margins white; stamens 18–20, filaments pale pink, white distally, anthers white; styles 2 or 3. |
Pomes | pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous; sepals flat, margins villous, villous; navel slightly open; style remnants 2/3 from base. |
purple-black, broadly ellipsoid to broadly obovoid, rarely cylindric, 9–11 × 8–11 mm, slightly shiny, not or slightly glaucous, villous; sepals flat or ascending, margins tomentose, strigose; forming star over open navel; style remnants 4/5 from base. |
Pyrenes | [1 or]2(or 3). |
2 or 3. |
2n | = 68 (Germany). |
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Cotoneaster fangianus |
Cotoneaster villosulus |
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Phenology | Flowering May; fruiting Oct–Nov. | Flowering May (often reflowering Sep); fruiting Aug–Nov. |
Habitat | Disturbed mesic forests | Thickets, open forests |
Elevation | 0–200 m (0–700 ft) | 0–50 m (0–200 ft) |
Distribution |
OR; Asia (China) [Introduced in North America; introduced also in Europe] |
WA; Asia (China) [Introduced in North America; introduced also in Europe] |
Discussion | Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The difficulties of typification and taxonomy for black-fruited species were discussed by J. Fryer and B. Hylmö (1997). Some collections of Cotoneaster villosulus from eastern Washington need study. They approach the very similar C. hsingshangensis J. Fryer & B. Hylmö, a Chinese species with larger, more wrinkled leaves, more globose pomes, and less densely hairy flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 467. | FNA vol. 9, p. 464. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. acutifolius var. villosulus | |
Name authority | T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) | (Rehder & E. H. Wilson) Flinck & B. Hylmö: Bot. Not. 115: 383. (1962) |
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