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Fang's cotoneaster

Mclaren's cotoneaster

Habit Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. Shrubs or trees, 4–6 m. Stems loosely erect, spreading, stiff; branches spiraled, maroon, minutely verrucose, initially pilose-strigose.
Leaves

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

deciduous;

petiole 4–6 mm, densely pilose-strigose;

blade on sterile shoots elliptic, (35–)50–80(–103) x (13–)17–37(–42) mm, chartaceous, margins flat, base cuneate, veins 6–10, superficial or lightly sunken, apex acute or obtuse, abaxial surfaces color not recorded, villose-strigose, adaxial brownish green, soon green, dull, coating not recorded, flat between lateral veins, glabrate;

fall leaves yellow.

Inflorescences

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

on fertile shoots 40–50 mm with 2–4 leaves, 10–25-flowered, lax.

Pedicels

2–5 mm, tomentose-pilose.

3–5 mm, densely pilose-strigose.

Flowers

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

10–12 mm diam.;

buds white;

hypanthium cupulate, villose-strigose;

sepals: margins villous, apex acute, surfaces sparsely villous;

petals spreading, white, with large hair tuft;

stamens (16–)20, filaments white, anthers black, styles 1 or 2.

Pomes

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

maroon to purple-black, obovoid to elliptic-obovoid, 9–12 × 9–10 mm, slightly shiny, glaucous, sometimes sparsely villous;

sepals flat, margins glabrous, sparsely villous;

navel slightly open;

style remnants 9/10 from base.

Pyrenes

[1 or]2(or 3).

1 or 2.

Cotoneaster fangianus

Cotoneaster transens

Phenology Flowering May; fruiting Oct–Nov. Flowering Jun; fruiting Oct–Nov.
Habitat Disturbed mesic forests Thickets
Elevation 0–200 m (0–700 ft) 0–50 m (0–200 ft)
Distribution
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
Discussion

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. Lingdi and A. R. Brach (2003) synonymized Cotoneaster transens with C. glaucophyllus Franchet (as C. glaucophyllus var. glaucophyllus). Cotoneaster transens is a deciduous species with purple-black pomes and is not closely related to C. glaucophyllus, an evergreen species with smaller leaves, smaller red pomes, and smaller flowers, among other differences.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 457.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. vestitus, C. villosulus, C. ×suecicus
Name authority T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) G. Klotz: Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 17(3): 337, fig. 4. (1968)
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