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Fang's cotoneaster

Himalayan cotoneaster, khasiberry cotoneaster, simons' cotoneaster

Habit Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. Shrubs, 2–4(–6) m; ± strict or ascending, crown narrow or ± columnar.
Stems

erect, stiff;

branches spiraled, divaricate, lateral branches ± straight, mostly suppressed, short, grayish brown, initially densely strigose.

Leaves

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

deciduous (sometimes tardily so on vigorous sprouts);

petiole 3–5 mm, strigose;

blade on vigorous shoots broadly elliptic to ovate, sometimes elliptic, rarely suborbiculate, 17–36 × 10–25 mm, chartaceous or subcoriaceous, base obtuse or cuneate, margins flat, veins 4 or 5, superficial or slightly sunken, apex acuminate or acute, abaxial surfaces light green, initially strigose, usually becoming sparsely so, adaxial green to dark green, shiny, not glaucous, flat or faintly bulging between lateral veins, or slightly rugose, sparsely strigose;

fall leaves yellow to bright orange or fiery red.

Inflorescences

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

on fertile shoots 15–30 mm with 4 leaves, 2–6-flowered, compact.

Pedicels

2–5 mm, tomentose-pilose.

2–5 mm, strigose.

Flowers

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

pendent, (5–)8–10 mm, open;

hypanthium campanulate, strigose;

sepals: margins villous, borders narrow, glabrous, apex recurved, long acuminate or cuspidate, surfaces strigose;

petals erect-incurved, dark red with white border;

stamens 20, filaments pink, anthers white;

styles (2 or)3–5.

Pomes

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

bright orange to orange-red, usually obovoid, rarely cylindric or broadly obovoid, 7–14 × 6–10 mm, shiny, not glaucous, glabrous or sparsely villous;

sepals erect or ascending, rarely nearly flat, margins villous, glabrous or sparsely villous;

navel open;

style remnants 2/3 from base.

Pyrenes

[1 or]2(or 3).

(2 or)3–5.

Cotoneaster fangianus

Cotoneaster simonsii

Phenology Flowering May; fruiting Oct–Nov. Flowering May–Jun; fruiting Oct–Apr.
Habitat Disturbed mesic forests Shores, clearings, disturbed or open forests, thickets
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
CA; OR; TN; WA; BC; Asia (Bhutan, India) [Introduced in North America; introduced also in Europe, Pacific Islands (New Zealand)]
[WildflowerSearch map]
[BONAP county map]
Discussion

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Reports of Cotoneaster simonsii from Ohio (M. A. Vincent and A. W. Cusick 1998; T. S. Cooperrider et al. 2001) are here referred to C. divaricatus. The binomial C. symondsii T. Moore was proposed as an earlier name for C. simonsii (A. Kumar and G. Panigrahi 1992; W. B. Dickoré and G. Kasperek 2010), but J. Fryer and P. F. Zika (2014) typified C. symondsii so that it is a synonym of the high Himalayan species C. marginatus Lindley ex Loudon.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 460.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Name authority T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) Baker: in W. W. Saunders, Refug. Bot. 1: plate 55. (1869)
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