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Fang's cotoneaster

cotoneaster, willow-leaf cotoneaster

Habit Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. Shrubs or trees, 1–8 m. Stems erect or arching [prostrate]; branches spiraled or distichous, arching, maroon, slender, initially tomentose-strigose.
Leaves

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

persistent;

petiole 3–8 mm, tomentose-strigose;

blade narrowly elliptic to elliptic-lanceolate, 30–90 × 8–23 mm, coriaceous, base cuneate, margins strongly revolute, veins 7–12, deeply sunken, apex gradually tapered, acute or acuminate, abaxial surfaces grayish green, initially densely villous, becoming glabrous or glabrate, adaxial dark green, shiny, not glaucous, bulging between lateral veins, sparsely pilose initially.

Inflorescences

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

on fertile shoots 20–60 mm with 3 or 4 leaves, 10–50-flowered, compact.

Pedicels

2–5 mm, tomentose-pilose.

0.5–3(–4 mm), densely strigose-villous.

Flowers

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

(5–)6–9 mm diam.;

buds white;

hypanthium funnelform, pilose;

sepals: margins sparsely villous, apex acute, surfaces sparsely pilose;

petals spreading, white, glabrous;

stamens 20, filaments white, anthers red-purple;

styles 2 or 3(or 4).

Pomes

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

bright red (or with hint of crimson), broadly obovoid to globose, (5–)6–8.5 x (5–)6–8 mm, shiny, not glaucous, sparsely pilose;

sepals depressed or flat, sparsely pilose;

navel slightly open;

style remnants at apex.

Pyrenes

[1 or]2(or 3).

2 or 3(or 4).

2n

= 34 (Germany).

Cotoneaster fangianus

Cotoneaster salicifolius

Phenology Flowering May; fruiting Oct–Nov. Flowering May–Jun; fruiting Oct–Apr.
Habitat Disturbed mesic forests Thickets, mossy cliffs, cracks in walls, lakeshores, urban waste ground
Elevation 0–200 m (0–700 ft) 0–200 m (0–700 ft)
Distribution
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
WA; BC; Asia (China: Sichuan, Xizang) [Introduced in North America; introduced also in Europe]
[BONAP county map]
Discussion

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. Lingdi and A. R. Brach (2003) included as varieties of Cotoneaster salicifolius several taxa that here are considered full species, such as C. henryanus (C. K. Schneider) Rehder & E. H. Wilson, C. hylmoei Flinck & J. Fryer, and C. rugosus E. Pritzel.

A horticultural hybrid, presumably between Cotoneaster dammeri and C. salicifolius, is known in gardens as the cultivar Cotoneaster 'Hybridus Pendulus' and was recently collected as a local escape from cultivation in King County, Washington. This hybrid would key to C. x\suecicus but differs in its limp branching, much larger leaves (20–80 × 10–30 mm) and more numerous and smaller diameter flowers (9–11 mm wide). The foliage of the hybrid closely resembles C. salicifolius, but it has fewer (4–13) and larger flowers and is less than 1 m in height.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 454.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Name authority T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) Franchet: Nouv. Arch. Mus. Hist. Nat., sér. 2, 8: 225. (1885)
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