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Fang's cotoneaster

herringbone cotoneaster, rock cotoneaster, rockspray cotoneaster, wall cotoneaster, wall or herringbone cotoneaster

Habit Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. Shrubs, 0.5–1 m. Stems ascending to erect, spreading horizontally; branches distichous, maroon, initially densely yellow-strigose.
Leaves

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

deciduous (tardily so on a few vigorous shoots);

petiole 2–3 mm, strigose;

blade elliptic or broadly elliptic, on vigorous shoots usually broadly elliptic, sometimes suborbiculate, orbiculate, or broadly obovate, 4–14 × 5–9 mm, subcoriaceous, base obtuse or cuneate, margins flat, veins 2–4, superficial, apex acute, apiculate or obtuse, mucronulate, abaxial surfaces green, sparsely long-strigose, adaxial dark green, shiny, not glaucous, flat between lateral veins, smooth to slightly rugose, glabrescent;

fall leaves orange to rich red.

Inflorescences

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

on fertile shoots 8–20 mm with 3 or 4 leaves, 1–3-flowered, compact.

Pedicels

2–5 mm, tomentose-pilose.

1–2 mm, strigose.

Flowers

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

5–7 mm, closed;

hypanthium funnelform, sparsely pilose-strigose;

sepals: margins villous, apex acute or acuminate, surfaces sparsely strigose;

petals erect-incurved, dark pink to dark red, base blackish red, margins pale crimson;

stamens (8–)10(–14), filaments dark red, anthers white;

styles 2 or 3(or 4).

Pomes

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

bright red, usually broadly obovoid, sometimes obovoid, rarely subglobose, 5–8.5 × 4.5–7.7 mm, shiny, not glaucous, glabrate;

sepals suberect, sparsely strigose or glabrate;

navel open;

style remnants 1/2–3/4 from base.

Pyrenes

[1 or]2(or 3).

2 or 3(or 4), distinct.

2n

= 68 (Germany).

Cotoneaster fangianus

Cotoneaster horizontalis

Phenology Flowering May; fruiting Oct–Nov. Flowering Apr–May; fruiting Sep–Mar.
Habitat Disturbed mesic forests Thickets, disturbed forests, paths, cliffs, pilings, rockwork
Elevation 0–200 m (0–700 ft) 0–500 m (0–1600 ft)
Distribution
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
CA; OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe]
[WildflowerSearch map]
[BONAP county map]
Discussion

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. Lingdi and A. R. Brach (2003) included within Cotoneaster horizontalis several taxa recognized here as distinct species, including C. flinckii J. Fryer & B. Hylmö and C. perpusillus (C. K. Schneider) Flinck & B. Hylmö. Reports of C. horizontalis from Ontario (J. K. Morton and J. M. Venn 1990) are here referred to C. divaricatus. Reports of C. horizontalis from Indiana are not supported by specimens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 462.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Name authority T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) Decaisne: Ann. Gén. Hort. 22: 168. (1877)
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