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Fang's cotoneaster

Gamble's cotoneaster

Habit Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. Shrubs or trees, 4–10 m. Stems loosely erect, spreading and arching; branches spiraled, brown or maroon-purple, initially tomentose-strigose.
Leaves

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

usually semipersistent (persisting to April if sheltered), sometimes deciduous;

petiole 4–10 mm, densely villose-strigose;

blade elliptic, (25–)50–127 × 13–49[–62 in cultivation] mm, chartaceous or subcoriaceous, base cuneate or obtuse, margins flat, veins 7 or 8, superficial, apex acute or obtuse, abaxial surfaces pale gray-green, initially villous, later glabrate, adaxial dark green, shiny, not glaucous, flat between lateral veins, sometimes becoming slightly rugose, glabrate;

fall leaves pale yellowish green.

Inflorescences

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

on fertile shoots 40–70 mm with 2 or 3 leaves, 5–34-flowered, lax.

Pedicels

2–5 mm, tomentose-pilose.

1–8 mm, sparsely villose-strigose.

Flowers

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

8–12.5 mm diam.;

buds white;

hypanthium cupulate, densely villous;

sepals: margins maroon, tomentose, apex acute, surfaces densely villous;

petals spreading, white, sometimes with hair tuft;

stamens 18–20, filaments white, anthers red-purple or dark red-purple;

styles (1 or)2.

Pomes

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

red, often tinged with pink, green, brown, or purple, obovoid, rarely subglobose, 8.5–12.8 × 8–12 mm, dull, slightly glaucous, sparsely villous;

sepals flat, margins tomentose, densely villous;

navel slightly open;

style remnants near apex.

Pyrenes

[1 or]2(or 3).

[1 or]2.

Cotoneaster fangianus

Cotoneaster gamblei

Phenology Flowering May; fruiting Oct–Nov. Flowering May; fruiting Sep–Nov.
Habitat Disturbed mesic forests Mesic thickets, flood plains, disturbed forests
Elevation 0–200 m (0–700 ft) 0–50 m (0–200 ft)
Distribution
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
WA; Asia (Bhutan, India) [Introduced in North America]
[BONAP county map]
Discussion

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 456.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Name authority T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 15: 530. (1966)
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