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Fang's cotoneaster

diel's cotoneaster

Habit Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. Shrubs, 2–4 m. Stems narrowly erect, spreading; branches distichous, grayish brown, initially tomentose-pilose.
Leaves

deciduous;

petiole 2–4 mm, tomentose;

blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose;

fall leaves yellowish.

deciduous;

petiole 2–5 mm, tomentose-pilose;

blade ovate to elliptic or broadly ovate, 9–38 × 7–23 mm, coriaceous or subcoriaceous, base obtuse or cuneate, margins flat, not revolute, veins 2–4, lightly sunken, apex acute or acuminate, abaxial surfaces densely grayish tomentose, adaxial dark green to grayish green, shiny, not glaucous, flat, rarely slightly bulging, between lateral veins, slightly rugose, pilose;

fall leaves bright yellow to orange or red.

Inflorescences

on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact.

on fertile shoots 10–35 mm with 4 leaves, 2–7(–10)-flowered, compact.

Pedicels

2–5 mm, tomentose-pilose.

1–3 mm, tomentose-strigose.

Flowers

erect, 4–6 mm;

hypanthium cupulate, densely villous;

sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally;

petals erect, pink or fading whitish;

stamens (16–)19 or 20, filaments pale pink or white, anthers white;

styles [1 or]2(or 3).

erect, 6–7.5 mm, closed;

hypanthium funnelform, proximally tomentose-strigose, distally tomentose-pilose;

sepals: margins reddish tomentose, borders purple, broad, glabrous, apex cuspidate or apiculate, surfaces pilose;

petals erect-incurved, dark pink to red, base dark red, margins off-white;

stamens 14–20, filaments red to dark red, anthers white;

styles 3 or 4(or 5).

Pomes

pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous;

sepals flat, margins villous, villous;

navel slightly open;

style remnants 2/3 from base.

bright red, broadly obovoid, sometimes globose or depressed-globose, 5–9.1 × 5.1–8.8 mm, slightly shiny, not glaucous, pilose;

sepals flat, tomentose;

navel open;

style remnants 1/2–2/3 from base.

Pyrenes

[1 or]2(or 3).

3 or 4(or 5).

Cotoneaster fangianus

Cotoneaster dielsianus

Phenology Flowering May; fruiting Oct–Nov. Flowering May–Jun; fruiting Aug–Mar.
Habitat Disturbed mesic forests Edges of woods, disturbed forests, flood plains, thickets, waste ground, cliffs, meadows, brushy wet prairie remnants
Elevation 0–200 m (0–700 ft) 0–500 m (0–1600 ft)
Distribution
from FNA
OR; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
Discussion

Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

L. Lingdi and A. R. Brach (2003) included within Cotoneaster dielsianus several taxa recognized here as distinct species, including C. bradyi J. Fryer & E. C. Nelson and C. splendens Flinck & B. Hylmö. Some collections from Oregon approach an undescribed species from China but differ in fruit shape.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 467. FNA vol. 9, p. 466.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Name authority T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) E. Pritz ex Diels: Bot. Jahrb. Syst. 29: 385. (1900)
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