Cotoneaster fangianus |
Cotoneaster cochleatus |
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Fang's cotoneaster |
Yunnan cotoneaster |
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Habit | Shrubs, 2–2.5 m. Stems erect, arching, spreading; branches distichous, maroon, purple, or brown, initially tomentose-pilose. | Shrubs, to 0.4 m. Stems prostrate or nearly so, carpeting, rooting; branches spiraled and distichous, dense, red to purple-black, slender, initially yellow-green strigose. |
Leaves | deciduous; petiole 2–4 mm, tomentose; blade elliptic, broadly elliptic, or ovate, rarely suborbiculate, 21–56 × 13–30 mm, subcoriaceous, base obtuse or truncate, margins flat, veins 4–6, slightly to deeply sunken, apex obtuse or acute, abaxial surfaces grayish, initially moderately to densely villous, adaxial dark green, dull, not glaucous, flat between lateral veins, slightly rugose, pilose; fall leaves yellowish. |
persistent; petiole 1–4 mm, strigose; blade obovate to broadly obovate, rarely suborbiculate, 5–14 × 3–9 mm, coriaceous, base obtuse or broadly cuneate, margins slightly revolute, veins 2 or 3, superficial, apex obtuse, sometimes emarginate, abaxial surfaces grayish, reticulate, initially densely strigose-villous, adaxial dark green, shiny, not glaucous, sometimes lightly rugose, glabrescent. |
Inflorescences | on fertile shoots 20–30 mm, usually with 2 or 3 leaves, 3–15-flowered, compact. |
on fertile shoots 8–15 mm, usually with 4 leaves, 1(–3)-flowered. |
Pedicels | 2–5 mm, tomentose-pilose. |
3–5 mm, strigose. |
Flowers | erect, 4–6 mm; hypanthium cupulate, densely villous; sepals: margins villous, apex acute, surfaces villous, sparsely hairy submarginally; petals erect, pink or fading whitish; stamens (16–)19 or 20, filaments pale pink or white, anthers white; styles [1 or]2(or 3). |
7–10 mm diam.; buds pinkish; hypanthium cupulate, strigose or pilose-strigose; sepals: margins villous, apex obtuse or acute, surfaces initially sparsely pilose-strigose; petals spreading, white, glabrous; stamens (15–)20, filaments white, anthers dark purple; styles 2(or 3). |
Pomes | pendent, bright to dark red, obovoid to obconic, 8–10 × 8–9 mm, apex nearly flat, reflectance and coating not recorded, succulent, villous; sepals flat, margins villous, villous; navel slightly open; style remnants 2/3 from base. |
bright red to crimson, subglobose, 7–9 × 8–10 mm, slightly shiny, not glaucous, sparsely pilose; sepals suberect, sparsely strigose; navel slightly open; style remnants at apex on small projection. |
Pyrenes | [1 or]2(or 3). |
2(or 3). |
2n | = 68 (Germany). |
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Cotoneaster fangianus |
Cotoneaster cochleatus |
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Phenology | Flowering May; fruiting Oct–Nov. | Flowering May–Jun; fruiting Sep–Nov. |
Habitat | Disturbed mesic forests | Forest edges |
Elevation | 0–200 m (0–700 ft) | 0–50 m (0–200 ft) |
Distribution |
OR; Asia (China) [Introduced in North America; introduced also in Europe] |
NY; Asia (China) [Introduced in North America; introduced also in Europe] |
Discussion | Reports of Cotoneaster mucronatus Franchet escaped in Oregon (P. F. Zika and E. R. Alverson 2005) are here referred to C. fangianus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cotoneaster cochleatus was treated as a variety of C. microphyllus by L. Lingdi and A. R. Brach (2003); here the two are distinguished at species rank, following H. Nybom et al. (2005). Plants of C. microphyllus have a suberect habit, usually elliptic leaves (rarely broadly obovate) with acute apices, and pomes 6 mm wide; C. cochleatus is always prostrate and has usually obovate leaves (rarely suborbiculate) with blunt apices, and pomes 8–10 mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 467. | FNA vol. 9, p. 459. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. buxifolius, C. microphyllus var. cochleatus, C. thymifolius var. cochleatus | |
Name authority | T. T. Yu: Acta Phytotax. Sin. 8: 219. (1963) | (Franchet) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 6: 952. (1957) |
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