Cotoneaster dielsianus |
Cotoneaster villosulus |
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diel's cotoneaster |
taiping cotoneaster |
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Habit | Shrubs, 2–4 m. Stems narrowly erect, spreading; branches distichous, grayish brown, initially tomentose-pilose. | Shrubs or trees, 3–5 m. Stems erect, arching, spreading, coarse; branches distichous or spiraled, brown, initially densely villose-strigose. |
Leaves | deciduous; petiole 2–5 mm, tomentose-pilose; blade ovate to elliptic or broadly ovate, 9–38 × 7–23 mm, coriaceous or subcoriaceous, base obtuse or cuneate, margins flat, not revolute, veins 2–4, lightly sunken, apex acute or acuminate, abaxial surfaces densely grayish tomentose, adaxial dark green to grayish green, shiny, not glaucous, flat, rarely slightly bulging, between lateral veins, slightly rugose, pilose; fall leaves bright yellow to orange or red. |
deciduous; petiole 2–5 mm, strigose; blade elliptic or ovate, 32–72(–104) x 18–37(–46) mm, subcoriaceous to chartaceous, base obtuse or cuneate, margins flat, not revolute, veins 5–8, lightly sunken, apex attenuate to long-acuminate, often acute on short shoots, abaxial surfaces green, shiny, sparsely pilose-strigose, adaxial dark reddish green, shiny, not glaucous, soon dark green, dullish, bulging between lateral veins, rugose, pilose-strigose; fall leaves purple and rich dark red. |
Inflorescences | on fertile shoots 10–35 mm with 4 leaves, 2–7(–10)-flowered, compact. |
on fertile shoots 25–50 mm with 4 leaves, 3–5(–7)-flowered, mostly compact. |
Pedicels | 1–3 mm, tomentose-strigose. |
3–7 mm, densely strigose. |
Flowers | erect, 6–7.5 mm, closed; hypanthium funnelform, proximally tomentose-strigose, distally tomentose-pilose; sepals: margins reddish tomentose, borders purple, broad, glabrous, apex cuspidate or apiculate, surfaces pilose; petals erect-incurved, dark pink to red, base dark red, margins off-white; stamens 14–20, filaments red to dark red, anthers white; styles 3 or 4(or 5). |
6–8 mm, opening small; hypanthium cupulate, tomentose-strigose; sepals: margins tomentose, borders brown, broad, glabrous, apex acute, obtuse, or apiculate, surfaces strigose; petals erect-incurved, pink or red, base dark red, margins white; stamens 18–20, filaments pale pink, white distally, anthers white; styles 2 or 3. |
Pomes | bright red, broadly obovoid, sometimes globose or depressed-globose, 5–9.1 × 5.1–8.8 mm, slightly shiny, not glaucous, pilose; sepals flat, tomentose; navel open; style remnants 1/2–2/3 from base. |
purple-black, broadly ellipsoid to broadly obovoid, rarely cylindric, 9–11 × 8–11 mm, slightly shiny, not or slightly glaucous, villous; sepals flat or ascending, margins tomentose, strigose; forming star over open navel; style remnants 4/5 from base. |
Pyrenes | 3 or 4(or 5). |
2 or 3. |
2n | = 68 (Germany). |
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Cotoneaster dielsianus |
Cotoneaster villosulus |
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Phenology | Flowering May–Jun; fruiting Aug–Mar. | Flowering May (often reflowering Sep); fruiting Aug–Nov. |
Habitat | Edges of woods, disturbed forests, flood plains, thickets, waste ground, cliffs, meadows, brushy wet prairie remnants | Thickets, open forests |
Elevation | 0–500 m (0–1600 ft) | 0–50 m (0–200 ft) |
Distribution |
OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe] |
WA; Asia (China) [Introduced in North America; introduced also in Europe] |
Discussion | L. Lingdi and A. R. Brach (2003) included within Cotoneaster dielsianus several taxa recognized here as distinct species, including C. bradyi J. Fryer & E. C. Nelson and C. splendens Flinck & B. Hylmö. Some collections from Oregon approach an undescribed species from China but differ in fruit shape. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The difficulties of typification and taxonomy for black-fruited species were discussed by J. Fryer and B. Hylmö (1997). Some collections of Cotoneaster villosulus from eastern Washington need study. They approach the very similar C. hsingshangensis J. Fryer & B. Hylmö, a Chinese species with larger, more wrinkled leaves, more globose pomes, and less densely hairy flowers. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 466. | FNA vol. 9, p. 464. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. acutifolius var. villosulus | |
Name authority | E. Pritz ex Diels: Bot. Jahrb. Syst. 29: 385. (1900) | (Rehder & E. H. Wilson) Flinck & B. Hylmö: Bot. Not. 115: 383. (1962) |
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