Cotoneaster dielsianus |
Cotoneaster divaricatus |
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diel's cotoneaster |
bearberry cotoneaster, spreading cotoneaster, stretch-fruit cotoneaster |
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Habit | Shrubs, 2–4 m. Stems narrowly erect, spreading; branches distichous, grayish brown, initially tomentose-pilose. | Shrubs, 1.5–2 m; crown spreading, rounded. |
Stems | erect; branches divaricate or spiraled, spreading, lateral branches well developed, usually arching, long, brownish violet, initially strigose. |
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Leaves | deciduous; petiole 2–5 mm, tomentose-pilose; blade ovate to elliptic or broadly ovate, 9–38 × 7–23 mm, coriaceous or subcoriaceous, base obtuse or cuneate, margins flat, not revolute, veins 2–4, lightly sunken, apex acute or acuminate, abaxial surfaces densely grayish tomentose, adaxial dark green to grayish green, shiny, not glaucous, flat, rarely slightly bulging, between lateral veins, slightly rugose, pilose; fall leaves bright yellow to orange or red. |
deciduous; petiole 3–4 mm, strigose; blade elliptic, rarely ovate or suborbiculate, on sterile shoots usually elliptic or broadly elliptic, 10–30 × 7–21 mm, chartaceous, base obtuse, margins flat, not revolute, sometimes undulate, veins 3 or 4, superficial or slightly sunken, apex acute, rarely acuminate, abaxial surfaces pale green, initially strigose, adaxial dark green, shiny, not glaucous, flat between lateral veins, initially sparsely strigose; fall leaves orange, red, or reddish purple. |
Inflorescences | on fertile shoots 10–35 mm with 4 leaves, 2–7(–10)-flowered, compact. |
on fertile shoots 10–20 mm with 4 leaves, 1–4-flowered, compact. |
Pedicels | 1–3 mm, tomentose-strigose. |
1–3 mm, strigose. |
Flowers | erect, 6–7.5 mm, closed; hypanthium funnelform, proximally tomentose-strigose, distally tomentose-pilose; sepals: margins reddish tomentose, borders purple, broad, glabrous, apex cuspidate or apiculate, surfaces pilose; petals erect-incurved, dark pink to red, base dark red, margins off-white; stamens 14–20, filaments red to dark red, anthers white; styles 3 or 4(or 5). |
6–8 mm, opening small; hypanthium cupulate, strigose; sepals: margins villous, apex acute or acuminate, surfaces strigose; petals erect-incurved, pale red, base dark red, border white; stamens 10–15, filaments dark red, whitish distally, anthers white; styles (1 or)2(or 3). |
Pomes | bright red, broadly obovoid, sometimes globose or depressed-globose, 5–9.1 × 5.1–8.8 mm, slightly shiny, not glaucous, pilose; sepals flat, tomentose; navel open; style remnants 1/2–2/3 from base. |
dark red to ruby (finally blackish red), cylindric, oblong-ellipsoid, or narrowly obovoid, 8–11 × 4–8 mm, shiny, not glaucous, sparsely pilose; sepals suberect, sparsely pilose; navel open; style remnants 2/3 from base. |
Pyrenes | 3 or 4(or 5). |
(1 or)2(or 3) [rarely 4]. |
Cotoneaster dielsianus |
Cotoneaster divaricatus |
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Phenology | Flowering May–Jun; fruiting Aug–Mar. | Flowering Apr–May; fruiting Sep–Dec. |
Habitat | Edges of woods, disturbed forests, flood plains, thickets, waste ground, cliffs, meadows, brushy wet prairie remnants | Thickets, edges, disturbed forests, flood plains, rarely epiphytic on maple |
Elevation | 0–500 m (0–1600 ft) | 0–700 m (0–2300 ft) |
Distribution |
OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe] |
IL; IN; KY; MA; MI; NY; OH; OR; PA; WA; WI; BC; ON; Asia (China) [Introduced in North America; introduced also in Europe]
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Discussion | L. Lingdi and A. R. Brach (2003) included within Cotoneaster dielsianus several taxa recognized here as distinct species, including C. bradyi J. Fryer & E. C. Nelson and C. splendens Flinck & B. Hylmö. Some collections from Oregon approach an undescribed species from China but differ in fruit shape. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Although rarely collected, Cotoneaster divaricatus is the most widely escaped cotoneaster in eastern North America. Misidentified specimens of it have been reported from New York as C. hupehensis Rehder & E. H. Wilson (R. S. Mitchell and G. C. Tucker 1997), from Illinois as C. acutifolius (J. T. Kartesz and C. A. Meacham 2003), from Indiana and Ontario as C. horizontalis (Kartesz and Meacham), from Kentucky as C. apiculatus Rehder & E. H. Wilson (Kartesz and Meacham), and from Wisconsin as C. lucidus (Kartesz and Meacham). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 466. | FNA vol. 9, p. 461. |
Parent taxa | ||
Sibling taxa | ||
Name authority | E. Pritz ex Diels: Bot. Jahrb. Syst. 29: 385. (1900) | Rehder & E. H. Wilson: in C. S. Sargent, Pl. Wilson. 1: 157. (1912) |
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