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necklace cotoneaster, winter-green cotoneaster

Gamble's cotoneaster

Habit Shrubs, 0.5–1[–2.5] m. Stems weakly ascending or mound-forming [prostrate]; branches spiraled or distichous, dense, maroon, initially strigose. Shrubs or trees, 4–10 m. Stems loosely erect, spreading and arching; branches spiraled, brown or maroon-purple, initially tomentose-strigose.
Leaves

persistent;

petiole 1–3 mm, pilose-strigose;

blade elliptic to narrowly elliptic, rarely lanceolate, 6–12(–20) x 2–6(–8) mm, subcoriaceous or coriaceous, base cuneate, margins revolute, veins 3–5, superficial, apex obtuse or acute, abaxial surfaces grayish green, reticulate, initially pilose-strigose, adaxial grayish green [rarely green], dull to slightly shiny, coating not recorded, slightly rugose, sparsely pilose or glabrous.

usually semipersistent (persisting to April if sheltered), sometimes deciduous;

petiole 4–10 mm, densely villose-strigose;

blade elliptic, (25–)50–127 × 13–49[–62 in cultivation] mm, chartaceous or subcoriaceous, base cuneate or obtuse, margins flat, veins 7 or 8, superficial, apex acute or obtuse, abaxial surfaces pale gray-green, initially villous, later glabrate, adaxial dark green, shiny, not glaucous, flat between lateral veins, sometimes becoming slightly rugose, glabrate;

fall leaves pale yellowish green.

Inflorescences

on fertile shoots 8–12 mm with 3 or 4 leaves, 1-flowered, 2–4-flowered on shoot tips.

on fertile shoots 40–70 mm with 2 or 3 leaves, 5–34-flowered, lax.

Pedicels

1–3 mm, pilose-strigose.

1–8 mm, sparsely villose-strigose.

Flowers

9–13 mm diam.;

buds pink;

hypanthium cupulate, pilose-strigose;

sepals: margins villous, borders mostly reddish, membranous, apex acute or obtuse, surfaces pilose-strigose;

petals spreading, white;

stamens 17–21, filaments white, anthers purple-black;

styles 2(or 3).

8–12.5 mm diam.;

buds white;

hypanthium cupulate, densely villous;

sepals: margins maroon, tomentose, apex acute, surfaces densely villous;

petals spreading, white, sometimes with hair tuft;

stamens 18–20, filaments white, anthers red-purple or dark red-purple;

styles (1 or)2.

Pomes

orange-red to red, depressed-globose, 8–10 × 9–11 mm, shiny, not glaucous, glabrous;

sepals suberect, pilose;

navel not recorded;

style remnants at elongated apex.

red, often tinged with pink, green, brown, or purple, obovoid, rarely subglobose, 8.5–12.8 × 8–12 mm, dull, slightly glaucous, sparsely villous;

sepals flat, margins tomentose, densely villous;

navel slightly open;

style remnants near apex.

Pyrenes

2(or 3).

[1 or]2.

2n

= 34 (Germany).

Cotoneaster conspicuus

Cotoneaster gamblei

Phenology Flowering Apr–May; fruiting Oct–May. Flowering May; fruiting Sep–Nov.
Habitat Brushy edges in urban areas Mesic thickets, flood plains, disturbed forests
Elevation 0–50 m (0–200 ft) 0–50 m (0–200 ft)
Distribution
from FNA
WA; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
WA; Asia (Bhutan, India) [Introduced in North America]
[BONAP county map]
Discussion

G. Klotz (1963) emended the diagnosis of Cotoneaster conspicuus to recognize erect plants as var. conspicuus and more decumbent plants as var. decorus. This variation in habit is not significant taxonomically and not unusual for an outcrossing diploid, so the emended description by Klotz of the species is superfluous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 457. FNA vol. 9, p. 456.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. cochleatus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. microphyllus var. conspicuus, C. conspicuus var. decorus, C. conspicuus var. nanus, C. nanus, C. permutatus
Name authority (Messel) Messel: J. Roy. Hort. Soc. 59: 303. (1934) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 15: 530. (1966)
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