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Yunnan cotoneaster

dark cotoneaster, dark-fruit cotoneaster

Habit Shrubs, to 0.4 m. Stems prostrate or nearly so, carpeting, rooting; branches spiraled and distichous, dense, red to purple-black, slender, initially yellow-green strigose. Shrubs, 2–2.5 m. Stems loosely erect; branches spiraled, yellow brown to red brown, lenticellate, shiny, initially densely pilose.
Leaves

persistent;

petiole 1–4 mm, strigose;

blade obovate to broadly obovate, rarely suborbiculate, 5–14 × 3–9 mm, coriaceous, base obtuse or broadly cuneate, margins slightly revolute, veins 2 or 3, superficial, apex obtuse, sometimes emarginate, abaxial surfaces grayish, reticulate, initially densely strigose-villous, adaxial dark green, shiny, not glaucous, sometimes lightly rugose, glabrescent.

deciduous;

petiole 4–7 mm, tomentose-villous;

blade elliptic to ovate, 33–45 × 20–32 mm, chartaceous, base rounded, margins flat, veins 5–7, superficial, apex acute or obtuse, abaxial surfaces densely silvery-pilose-villous, adaxial green to dark green, dull to slightly shiny, not glaucous, flat between lateral veins, rugose, sparsely pilose;

fall leaves lacking notable color.

Inflorescences

on fertile shoots 8–15 mm, usually with 4 leaves, 1(–3)-flowered.

on fertile shoots 25–30 mm with 3 or 4 leaves, 5–13-flowered, pendent, lax.

Pedicels

3–5 mm, strigose.

3–8 mm, sometimes sparsely villous.

Flowers

7–10 mm diam.;

buds pinkish;

hypanthium cupulate, strigose or pilose-strigose;

sepals: margins villous, apex obtuse or acute, surfaces initially sparsely pilose-strigose;

petals spreading, white, glabrous;

stamens (15–)20, filaments white, anthers dark purple;

styles 2(or 3).

erect, 6–7 mm, open;

hypanthium cupulate, dark reddish brown, glabrous;

sepals: margins erose, apex acute or obtuse, surfaces glabrous, often apically villous;

petals erect, greenish white with pink, red, base slightly darkened, margins white, glabrous;

stamens 20(–22), filaments white, anthers white;

styles 2–4.

Pomes

bright red to crimson, subglobose, 7–9 × 8–10 mm, slightly shiny, not glaucous, sparsely pilose;

sepals suberect, sparsely strigose;

navel slightly open;

style remnants at apex on small projection.

purple-black, obovoid or globose, 7–9 × 7–9 mm, dull, glaucous with blue tinge, glabrous;

sepals suberect, glabrous or apically villous;

navel open;

style remnants 2/3 from base.

Pyrenes

2(or 3).

2–4.

2n

= 68 (Germany).

= 48–52, 68 (Russia).

Cotoneaster cochleatus

Cotoneaster melanocarpus

Phenology Flowering May–Jun; fruiting Sep–Nov. Flowering Apr–May; fruiting Jul–Aug.
Habitat Forest edges Forested ravines
Elevation 0–50 m (0–200 ft) 300–500 m (1000–1600 ft)
Distribution
from FNA
NY; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
MB; Eurasia (Russia, Ukraine) [Introduced in North America]
[BONAP county map]
Discussion

Cotoneaster cochleatus was treated as a variety of C. microphyllus by L. Lingdi and A. R. Brach (2003); here the two are distinguished at species rank, following H. Nybom et al. (2005). Plants of C. microphyllus have a suberect habit, usually elliptic leaves (rarely broadly obovate) with acute apices, and pomes 6 mm wide; C. cochleatus is always prostrate and has usually obovate leaves (rarely suborbiculate) with blunt apices, and pomes 8–10 mm wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cotoneaster melanocarpus is said to hybridize spontaneously with Sorbus aucuparia in Siberia, forming x\Sorbocotoneaster Pojarkova. The actual Cotoneaster parent is more likely to be a diploid and needs investigation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 459. FNA vol. 9, p. 467.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. buxifolius, C. microphyllus var. cochleatus, C. thymifolius var. cochleatus C. vulgaris var. melanocarpus
Name authority (Franchet) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 6: 952. (1957) (Ledebour) Loddiges: Fam. Nat. Syn. Monogr. 3: 223. (1847)
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