Cotoneaster cochleatus |
Cotoneaster lacteus |
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Yunnan cotoneaster |
cream-flower cotoneaster, cream-flower or late or parney's cotoneaster, late cotoneaster, milk-flower cotoneaster |
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Habit | Shrubs, to 0.4 m. Stems prostrate or nearly so, carpeting, rooting; branches spiraled and distichous, dense, red to purple-black, slender, initially yellow-green strigose. | Shrubs or trees, 2–9 m. Stems erect, arching, spreading, slender; branches spiraled or distichous, purple-black, initially yellowish tomentose-villous. |
Leaves | persistent; petiole 1–4 mm, strigose; blade obovate to broadly obovate, rarely suborbiculate, 5–14 × 3–9 mm, coriaceous, base obtuse or broadly cuneate, margins slightly revolute, veins 2 or 3, superficial, apex obtuse, sometimes emarginate, abaxial surfaces grayish, reticulate, initially densely strigose-villous, adaxial dark green, shiny, not glaucous, sometimes lightly rugose, glabrescent. |
persistent; petiole 5–8 mm, initially tomentose; blade obovate to elliptic, rarely broadly elliptic, (26–)40–85(–120) x (13–)19–45(–60) mm, coriaceous, base cuneate, margins flat or slightly revolute, veins 7–9, deeply sunken, apex short-tapered, acute, acuminate, or obtuse, abaxial surfaces initially yellowish tomentose-villous, adaxial dark green, dull or slightly shiny, not glaucous, bulging slightly between lateral veins, initially sparsely pilose. |
Inflorescences | on fertile shoots 8–15 mm, usually with 4 leaves, 1(–3)-flowered. |
on fertile shoots 45–100 mm with 4 leaves, 20–150-flowered, lax. |
Pedicels | 3–5 mm, strigose. |
1–5 mm, tomentose-strigose. |
Flowers | 7–10 mm diam.; buds pinkish; hypanthium cupulate, strigose or pilose-strigose; sepals: margins villous, apex obtuse or acute, surfaces initially sparsely pilose-strigose; petals spreading, white, glabrous; stamens (15–)20, filaments white, anthers dark purple; styles 2(or 3). |
6–9 mm diam.; buds creamy white, sometimes slightly pinkish; hypanthium funnelform, tomentose-strigose; sepals: margins villous, apex cuspidate, surfaces tomentose; petals spreading, creamy white, sometimes with hair tuft; stamens 20, filaments white, anthers red-purple; styles 2. |
Pomes | bright red to crimson, subglobose, 7–9 × 8–10 mm, slightly shiny, not glaucous, sparsely pilose; sepals suberect, sparsely strigose; navel slightly open; style remnants at apex on small projection. |
crimson, broadly obovoid, subglobose, or globose, 6.5–8.6 × 6.5–7.6 mm, shiny, not glaucous, sparsely villous; sepals depressed or slightly ascending, tomentose; navel slightly open or obscured by sepals; style remnants near apex. |
Pyrenes | 2(or 3). |
2. |
2n | = 68 (Germany). |
= 68 (Germany). |
Cotoneaster cochleatus |
Cotoneaster lacteus |
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Phenology | Flowering May–Jun; fruiting Sep–Nov. | Flowering May–Jul; fruiting Nov–Apr. |
Habitat | Forest edges | Thickets, disturbed forests, edges, urban waste ground, stonework, grassy bluffs, canyons, pond shores |
Elevation | 0–50 m (0–200 ft) | 0–700 m (0–2300 ft) |
Distribution |
NY; Asia (China) [Introduced in North America; introduced also in Europe] |
CA; OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe, Pacific Islands (New Zealand)]
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Discussion | Cotoneaster cochleatus was treated as a variety of C. microphyllus by L. Lingdi and A. R. Brach (2003); here the two are distinguished at species rank, following H. Nybom et al. (2005). Plants of C. microphyllus have a suberect habit, usually elliptic leaves (rarely broadly obovate) with acute apices, and pomes 6 mm wide; C. cochleatus is always prostrate and has usually obovate leaves (rarely suborbiculate) with blunt apices, and pomes 8–10 mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cotoneaster lacteus sometimes is synonymized with C. coriaceus Franchet (for example, L. Lingdi and A. R. Brach 2003). As treated here, plants of this species are identified by their fertile shoots to 10 cm, scarcely shiny leaves to 120 × 60 mm with adaxial surfaces bulging between lateral veins, flowers to 150 in lax inflorescences, 6–9 mm diam. with cream petals, and larger, later-ripening fruits. Plants of C. coriaceus have fertile shoots to 5 cm, shiny leaves to 52 × 24 mm with flat adaxial surfaces, flowers to 50 in compact inflorescences, 9–10 mm diam. with white petals, and smaller, earlier-ripening fruits. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 459. | FNA vol. 9, p. 454. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. buxifolius, C. microphyllus var. cochleatus, C. thymifolius var. cochleatus | |
Name authority | (Franchet) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 6: 952. (1957) | W. W. Smith: Notes Roy. Bot. Gard. Edinburgh 10: 23. (1917) |
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