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Yunnan cotoneaster

Gamble's cotoneaster

Habit Shrubs, to 0.4 m. Stems prostrate or nearly so, carpeting, rooting; branches spiraled and distichous, dense, red to purple-black, slender, initially yellow-green strigose. Shrubs or trees, 4–10 m. Stems loosely erect, spreading and arching; branches spiraled, brown or maroon-purple, initially tomentose-strigose.
Leaves

persistent;

petiole 1–4 mm, strigose;

blade obovate to broadly obovate, rarely suborbiculate, 5–14 × 3–9 mm, coriaceous, base obtuse or broadly cuneate, margins slightly revolute, veins 2 or 3, superficial, apex obtuse, sometimes emarginate, abaxial surfaces grayish, reticulate, initially densely strigose-villous, adaxial dark green, shiny, not glaucous, sometimes lightly rugose, glabrescent.

usually semipersistent (persisting to April if sheltered), sometimes deciduous;

petiole 4–10 mm, densely villose-strigose;

blade elliptic, (25–)50–127 × 13–49[–62 in cultivation] mm, chartaceous or subcoriaceous, base cuneate or obtuse, margins flat, veins 7 or 8, superficial, apex acute or obtuse, abaxial surfaces pale gray-green, initially villous, later glabrate, adaxial dark green, shiny, not glaucous, flat between lateral veins, sometimes becoming slightly rugose, glabrate;

fall leaves pale yellowish green.

Inflorescences

on fertile shoots 8–15 mm, usually with 4 leaves, 1(–3)-flowered.

on fertile shoots 40–70 mm with 2 or 3 leaves, 5–34-flowered, lax.

Pedicels

3–5 mm, strigose.

1–8 mm, sparsely villose-strigose.

Flowers

7–10 mm diam.;

buds pinkish;

hypanthium cupulate, strigose or pilose-strigose;

sepals: margins villous, apex obtuse or acute, surfaces initially sparsely pilose-strigose;

petals spreading, white, glabrous;

stamens (15–)20, filaments white, anthers dark purple;

styles 2(or 3).

8–12.5 mm diam.;

buds white;

hypanthium cupulate, densely villous;

sepals: margins maroon, tomentose, apex acute, surfaces densely villous;

petals spreading, white, sometimes with hair tuft;

stamens 18–20, filaments white, anthers red-purple or dark red-purple;

styles (1 or)2.

Pomes

bright red to crimson, subglobose, 7–9 × 8–10 mm, slightly shiny, not glaucous, sparsely pilose;

sepals suberect, sparsely strigose;

navel slightly open;

style remnants at apex on small projection.

red, often tinged with pink, green, brown, or purple, obovoid, rarely subglobose, 8.5–12.8 × 8–12 mm, dull, slightly glaucous, sparsely villous;

sepals flat, margins tomentose, densely villous;

navel slightly open;

style remnants near apex.

Pyrenes

2(or 3).

[1 or]2.

2n

= 68 (Germany).

Cotoneaster cochleatus

Cotoneaster gamblei

Phenology Flowering May–Jun; fruiting Sep–Nov. Flowering May; fruiting Sep–Nov.
Habitat Forest edges Mesic thickets, flood plains, disturbed forests
Elevation 0–50 m (0–200 ft) 0–50 m (0–200 ft)
Distribution
from FNA
NY; Asia (China) [Introduced in North America; introduced also in Europe]
[BONAP county map]
from FNA
WA; Asia (Bhutan, India) [Introduced in North America]
[BONAP county map]
Discussion

Cotoneaster cochleatus was treated as a variety of C. microphyllus by L. Lingdi and A. R. Brach (2003); here the two are distinguished at species rank, following H. Nybom et al. (2005). Plants of C. microphyllus have a suberect habit, usually elliptic leaves (rarely broadly obovate) with acute apices, and pomes 6 mm wide; C. cochleatus is always prostrate and has usually obovate leaves (rarely suborbiculate) with blunt apices, and pomes 8–10 mm wide.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 9, p. 459. FNA vol. 9, p. 456.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster Rosaceae > subfam. Amygdaloideae > tribe Maleae > Cotoneaster
Sibling taxa
C. adpressus, C. atropurpureus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. gamblei, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
C. adpressus, C. atropurpureus, C. cochleatus, C. conspicuus, C. crispii, C. dammeri, C. dielsianus, C. divaricatus, C. fangianus, C. franchetii, C. frigidus, C. hjelmqvistii, C. hodjingensis, C. horizontalis, C. integrifolius, C. lacteus, C. lucidus, C. magnificus, C. melanocarpus, C. miniatus, C. monopyrenus, C. nitens, C. pannosus, C. qungbixiensis, C. rehderi, C. salicifolius, C. simonsii, C. sternianus, C. tengyuehensis, C. transens, C. vestitus, C. villosulus, C. ×suecicus
Synonyms C. buxifolius, C. microphyllus var. cochleatus, C. thymifolius var. cochleatus
Name authority (Franchet) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 6: 952. (1957) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 15: 530. (1966)
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