Cotoneaster cochleatus |
Cotoneaster gamblei |
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Yunnan cotoneaster |
Gamble's cotoneaster |
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Habit | Shrubs, to 0.4 m. Stems prostrate or nearly so, carpeting, rooting; branches spiraled and distichous, dense, red to purple-black, slender, initially yellow-green strigose. | Shrubs or trees, 4–10 m. Stems loosely erect, spreading and arching; branches spiraled, brown or maroon-purple, initially tomentose-strigose. |
Leaves | persistent; petiole 1–4 mm, strigose; blade obovate to broadly obovate, rarely suborbiculate, 5–14 × 3–9 mm, coriaceous, base obtuse or broadly cuneate, margins slightly revolute, veins 2 or 3, superficial, apex obtuse, sometimes emarginate, abaxial surfaces grayish, reticulate, initially densely strigose-villous, adaxial dark green, shiny, not glaucous, sometimes lightly rugose, glabrescent. |
usually semipersistent (persisting to April if sheltered), sometimes deciduous; petiole 4–10 mm, densely villose-strigose; blade elliptic, (25–)50–127 × 13–49[–62 in cultivation] mm, chartaceous or subcoriaceous, base cuneate or obtuse, margins flat, veins 7 or 8, superficial, apex acute or obtuse, abaxial surfaces pale gray-green, initially villous, later glabrate, adaxial dark green, shiny, not glaucous, flat between lateral veins, sometimes becoming slightly rugose, glabrate; fall leaves pale yellowish green. |
Inflorescences | on fertile shoots 8–15 mm, usually with 4 leaves, 1(–3)-flowered. |
on fertile shoots 40–70 mm with 2 or 3 leaves, 5–34-flowered, lax. |
Pedicels | 3–5 mm, strigose. |
1–8 mm, sparsely villose-strigose. |
Flowers | 7–10 mm diam.; buds pinkish; hypanthium cupulate, strigose or pilose-strigose; sepals: margins villous, apex obtuse or acute, surfaces initially sparsely pilose-strigose; petals spreading, white, glabrous; stamens (15–)20, filaments white, anthers dark purple; styles 2(or 3). |
8–12.5 mm diam.; buds white; hypanthium cupulate, densely villous; sepals: margins maroon, tomentose, apex acute, surfaces densely villous; petals spreading, white, sometimes with hair tuft; stamens 18–20, filaments white, anthers red-purple or dark red-purple; styles (1 or)2. |
Pomes | bright red to crimson, subglobose, 7–9 × 8–10 mm, slightly shiny, not glaucous, sparsely pilose; sepals suberect, sparsely strigose; navel slightly open; style remnants at apex on small projection. |
red, often tinged with pink, green, brown, or purple, obovoid, rarely subglobose, 8.5–12.8 × 8–12 mm, dull, slightly glaucous, sparsely villous; sepals flat, margins tomentose, densely villous; navel slightly open; style remnants near apex. |
Pyrenes | 2(or 3). |
[1 or]2. |
2n | = 68 (Germany). |
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Cotoneaster cochleatus |
Cotoneaster gamblei |
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Phenology | Flowering May–Jun; fruiting Sep–Nov. | Flowering May; fruiting Sep–Nov. |
Habitat | Forest edges | Mesic thickets, flood plains, disturbed forests |
Elevation | 0–50 m (0–200 ft) | 0–50 m (0–200 ft) |
Distribution |
NY; Asia (China) [Introduced in North America; introduced also in Europe] |
WA; Asia (Bhutan, India) [Introduced in North America] |
Discussion | Cotoneaster cochleatus was treated as a variety of C. microphyllus by L. Lingdi and A. R. Brach (2003); here the two are distinguished at species rank, following H. Nybom et al. (2005). Plants of C. microphyllus have a suberect habit, usually elliptic leaves (rarely broadly obovate) with acute apices, and pomes 6 mm wide; C. cochleatus is always prostrate and has usually obovate leaves (rarely suborbiculate) with blunt apices, and pomes 8–10 mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 9, p. 459. | FNA vol. 9, p. 456. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. buxifolius, C. microphyllus var. cochleatus, C. thymifolius var. cochleatus | |
Name authority | (Franchet) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 6: 952. (1957) | G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 15: 530. (1966) |
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