Cotoneaster cochleatus |
Cotoneaster franchetii |
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Yunnan cotoneaster |
cotoneaster, Franchet's cotoneaster, orange cotoneaster |
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Habit | Shrubs, to 0.4 m. Stems prostrate or nearly so, carpeting, rooting; branches spiraled and distichous, dense, red to purple-black, slender, initially yellow-green strigose. | Shrubs, 2–3 m. Stems erect, arching, lax, slender; branches distichous, maroon, initially densely pilose-strigose. |
Leaves | persistent; petiole 1–4 mm, strigose; blade obovate to broadly obovate, rarely suborbiculate, 5–14 × 3–9 mm, coriaceous, base obtuse or broadly cuneate, margins slightly revolute, veins 2 or 3, superficial, apex obtuse, sometimes emarginate, abaxial surfaces grayish, reticulate, initially densely strigose-villous, adaxial dark green, shiny, not glaucous, sometimes lightly rugose, glabrescent. |
persistent, often 30–40 mm apart; petiole 2–4 mm, tomentose-pilose; blade ovate or elliptic, 22–58 × 12–29 mm, coriaceous, base cuneate or obtuse, margins flat, veins 4 or 5, sunken, apex acute or acuminate, mucronate, abaxial surfaces silvery-tomentose, adaxial grayish green, slightly shiny, not glaucous, flat, rarely slightly bulging between lateral veins, drying slightly rugose, pilose. |
Inflorescences | on fertile shoots 8–15 mm, usually with 4 leaves, 1(–3)-flowered. |
on fertile shoots 30–60 mm with 4 leaves, 5–15(–25)-flowered, compact. |
Pedicels | 3–5 mm, strigose. |
1–4 mm, densely long-silky hairy. |
Flowers | 7–10 mm diam.; buds pinkish; hypanthium cupulate, strigose or pilose-strigose; sepals: margins villous, apex obtuse or acute, surfaces initially sparsely pilose-strigose; petals spreading, white, glabrous; stamens (15–)20, filaments white, anthers dark purple; styles 2(or 3). |
(5–)7–8(–10) mm, opening small; hypanthium cupulate, densely long-silky hairy; sepals: margins villous, borders purple, glabrous, apex cuspidate or acuminate, surfaces silky hairy; petals erect-incurved, pink or red, base dark pink to dark red, margins off-white; stamens 16–21, filaments pink, distally white, anthers pink or purplish pink; styles 2 or 3(or 4). |
Pomes | bright red to crimson, subglobose, 7–9 × 8–10 mm, slightly shiny, not glaucous, sparsely pilose; sepals suberect, sparsely strigose; navel slightly open; style remnants at apex on small projection. |
orange-red, obovoid base rounded or obconic base cuneate, straight-sided, 8–12 × 6–9 mm, apex flat, shiny, not glaucous, sparsely pilose; sepals flat, often asymmetric with 2 sepals longer than others, tomentose; navel slightly open; style remnants 3/4 from base. |
Pyrenes | 2(or 3). |
2 or 3(or 4). |
2n | = 68 (Germany). |
= 68 (Germany). |
Cotoneaster cochleatus |
Cotoneaster franchetii |
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Phenology | Flowering May–Jun; fruiting Sep–Nov. | Flowering May–Jul; fruiting Oct–Apr. |
Habitat | Forest edges | Edges, thickets, disturbed forests, flood plains, urban waste ground, mossy ledges, wet prairie remnants |
Elevation | 0–50 m (0–200 ft) | 0–700 m (0–2300 ft) |
Distribution |
NY; Asia (China) [Introduced in North America; introduced also in Europe] |
CA; OR; WA; BC; Asia (China) [Introduced in North America; introduced also in Europe, Pacific Islands (New Zealand), Australia]
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Discussion | Cotoneaster cochleatus was treated as a variety of C. microphyllus by L. Lingdi and A. R. Brach (2003); here the two are distinguished at species rank, following H. Nybom et al. (2005). Plants of C. microphyllus have a suberect habit, usually elliptic leaves (rarely broadly obovate) with acute apices, and pomes 6 mm wide; C. cochleatus is always prostrate and has usually obovate leaves (rarely suborbiculate) with blunt apices, and pomes 8–10 mm wide. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Reports of Cotoneaster franchetii from Colorado (J. Ells 2006) are here referred to C. lucidus. A few specimens escaped in Washington have red filaments, not pink, but otherwise agree with C. franchetii. Other escapes in Washington suggest the closely related C. wardii W. W. Smith, distinguished by semievergreen habit, longer fertile shoots, more lax and elongate inflorescences, and white anthers. The entire complex needs biosystematic review. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 9, p. 459. | FNA vol. 9, p. 465. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. buxifolius, C. microphyllus var. cochleatus, C. thymifolius var. cochleatus | |
Name authority | (Franchet) G. Klotz: Wiss. Z. Martin-Luther-Univ. Halle-Wittenberg, Math.-Naturwiss. Reihe 6: 952. (1957) | Bois: Rev. Hort. 74: 380, figs. 159–161, 164. (1902) |
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