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cob cactus, white-column foxtail cactus

Nellie's pincushion cactus

Habit Plants usually branched and small stemmed (to 50 branches), sometimes unbranched and large stemmed, corncob-like or pinecone-like on below-ground portion, on above-ground portion only on oldest plants, distal portion of stem ± obscured by spines. Plants usually unbranched (or branches 2–3; branched profusely in cultivation), smooth.
Roots

± diffuse.

diffuse or short taproots, largest roots basally 19–63 (average 33)% of greater stem diam.

Stems

ovoid to cylindric (spheric), 4–16 × (2.2–)3–6(–6.5) cm;

tubercles (6–)8–11 × 3–6 mm, firm;

areolar glands absent;

parenchyma not mucilaginous;

druses in pith and cortex nearly microscopic, mostly spheric;

pith 1/8–1/4 of lesser stem diam.;

medullary vascular system absent.

deep-seated with only hemispheric apex protruding above ground level, spheric to cylindric, 1–2.7 × 0.6–1.7(–2.5) cm;

tubercles 2–5 × 2–4 mm;

areolar glands absent;

parenchyma not mucilaginous;

druses present in the stem and tubercle cortex (large lenticular druses absent);

pith 1/5–1/3 of lesser stem diam.;

medullary vascular system absent.

Spines

(17–)21–41 per areole, ashy white, gray, or pale tan, tips of largest spines pinkish tan to reddish brown or reddish black, all straight;

radial spines 15–41 per areole, gray, (5–)7–12(–13.5) mm, largest spines 0.1–0.2 mm diam.;

subcentral spines 0–6 per areole;

central spines usually 5;

outer central spines (1–)3(–7) per areole, erect or ascending;

inner central spines (0–)2 per areole, porrect or descending, longest spines (5–)10–15(–18) × 0.2–0.3(–0.5) mm.

15–28 per areole, appressed, lightly pigmented, pale tan to pinkish gray, weathering to gray, tips darker on some of largest spines, dark purplish brown to orange-brown, often partly overlain by an ephemeral whitish layer, peglike, not sharply needlelike, proximally compressed laterally;

radial spines 13–24 per areole, 3.5–5 × 0.1–0.5 mm, radial/central distinction obscure;

subcentral spines usually 1–4 in adaxial parts of areoles;

outer central spines (1–)3(–4) per areole, in bird’s-foot arrangement at adaxial parts of clusters in adults, straight, inner central spines 0 per areole, longest spines 4–6 × 0.3–0.6 mm.

Flowers

apical or nearly so, (18–)20–30(–32) × 20–45(–4) mm, sterile distal part of flower tube 5–8.5(–11) mm, longer than stamen-bearing part;

outer tepals conspicuously fringed;

inner tepals 21 per flower, pure white, pale rose-pink, or pale lavender-pink, darker centrally, midstripes ± inconspicuous, (9–)11–19 × 1.5–2.5(–3.5) mm;

outer filaments cream;

anthers pale yellow or nearly white;

stigma lobes 4–6(–8), white, (1.8–)2–4 mm.

nearly apical, 13–16 × 15–27 mm;

outer tepals fringed;

inner tepals 15 or 16 per flower, pale rose-pink to magenta, shading gradually to paler pink or white basally, sometimes with a darker midstripes distally but lacking well-defined midstripes, proximally whitish to pale yellow-green, 7–12 × 2.5–4 mm;

outer filaments whitish to pale yellow-green;

anthers yellow to orange-yellow;

stigma lobes 4–8, green, 0.5–1.5 mm.

Fruits

bright red [green to maroon], ellipsoid, cylindric, or narrowly obovoid, (8–)13–25 × 3.5–6.5(–7.5) mm, not very succulent;

floral remnant strongly persistent.

green (sometimes slightly yellow tinted), spheric, obovoid, or obconic, 1.5–6 × 1.5–4 mm, nearly dry;

floral remnant persistent but easily broken off.

Seeds

reddish brown, darker with age, obliquely obovoid, 0.9–1 mm, pitted.

black, ± obovoid (to weakly pyriform), strophiole squarish, 0.8–1 mm, glossy, deeply pitted.

2n

= 22 (as C. strobiliformis, C. varicolor, and Escobaria tuberculosa).

= 22.

Coryphantha tuberculosa

Coryphantha minima

Phenology Flowering (Apr-)May–Aug; fruiting Jul–Oct. Flowering spring (Mar-)Apr–May(-Jun); fruiting late summer–fall.
Habitat Stony grasslands, oak-juniper savannas, Larrea scrub, often with Agave lechuguilla, limestone mountainsides or igneous rocks and novaculite Grasslands, closely associated with mats of Selaginella arizonica, novaculite ridges
Elevation 500-1800(-2200) m (1600-5900(-7200) ft) 1200-1400 m (3900-4600 ft)
Distribution
from FNA
NM; TX; Mexico (Chihuahua, Coahuila, Durango)
from FNA
TX
Discussion

The names Coryphantha strobiliformis and Escobaria strobiliformis have been misapplied to C. tuberculosa by some recent authors (e.g., L. D. Benson 1982). Those names were based on Echinocactus strobiliformis Poselger, which is C. chihuahuensis (Britton & Rose) A. Berger.

Despite strong superficial similarity to other species in the genus, Coryphantha tuberculosa seems taxonomically isolated. Coryphantha tuberculosa superficially resembles C. sneedii, from which it is distinguished by (1) giant lenticular druses absent (abundant in older pith and cortex of C. sneedii); (2) fruits in region of sympatry always red (green in most U.S. populations of C. sneedii); (3) maximal expansion of flowers in late afternoon, sometimes remaining fully open at sunset (unlike any other species of Coryphantha); (4) flowers larger than those of C. sneedii, either pure white or a characteristic shade of pale lavender-pink, identifiable at a glance when flowers are alive and open; (5) anthers pale yellow, nearly white (bright yellow in C. sneedii); and (6) sterile distal part of receptacular tube longer than the stamen-bearing portion (short in C. sneedii).

On igneous and metamorphic substrates populations of Coryphantha tuberculosa mostly have unbranched stems. D. Weniger (1984) considered such populations to represent C. varicolor Tiegel; their reproductive structures, however, are identical to those of C. tuberculosa. Without seeds or flowers, mature specimens from such populations sometimes are indistinguishable from C. dasyacantha [hence the synonym C. dasyacantha var. varicolor (Tiegel) L. D. Benson].

Coryphantha tuberculosa is the type species of the segregate genus Escobaria, which includes the coryphanthas with pitted seeds.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Coryphantha minima is a dwarf cactus known only from the vicinity of Marathon, Texas, where it occurs among Selaginella in crevices of Caballos novaculite ledges. The characteristic spines appear blunt—almost club-shaped—and lie flat against the stem, making it one of the safest coryphanthas to handle. The tiny stems, large flowers, and unusual spines make this plant attractive to cactus fanciers. Its population, however, is extremely restricted.

Coryphantha minima is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4. FNA vol. 4, p. 233.
Parent taxa Cactaceae > subfam. Cactoideae > Coryphantha Cactaceae > subfam. Cactoideae > Coryphantha
Sibling taxa
C. alversonii, C. chaffeyi, C. chlorantha, C. dasyacantha, C. duncanii, C. echinus, C. hesteri, C. macromeris, C. minima, C. missouriensis, C. nickelsiae, C. ramillosa, C. recurvata, C. robbinsorum, C. robertii, C. robustispina, C. sneedii, C. sulcata, C. vivipara
C. alversonii, C. chaffeyi, C. chlorantha, C. dasyacantha, C. duncanii, C. echinus, C. hesteri, C. macromeris, C. missouriensis, C. nickelsiae, C. ramillosa, C. recurvata, C. robbinsorum, C. robertii, C. robustispina, C. sneedii, C. sulcata, C. tuberculosa, C. vivipara
Synonyms Mamillaria tuberculosa, C. dasyacantha var. varicolor, C. varicolor, Escobaria dasyacantha var. varicolor, Escobaria tuberculosa C. nellieae, Escobaria minima, Escobaria nellieae, Mammillaria nellieae
Name authority (Engelmann) A. Berger: Kakteen, 280. (1929) Baird: Amer. Bot. (Binghamton) 37: 150. (1931)
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