Cornus unalaschkensis |
Cornus racemosa |
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alaskan bunchberry, western bunchberry, western cordilleran bunchberry |
cornouiller à grappes, gray dogwood, northern swamp dogwood, panicle dogwood |
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Habit | Shrubs, to 5 m, flowering at 0.7 m; rhizomes present. | |
Stems | erect, green, 6–22 cm, appressed-hairy; nodes 4–6, internodes progressively longer distally; branches only at distalmost node, much shorter than distal internodes so stems appear unbranched. |
solitary, 2–10 dm apart; bark gray, brittle, verrucose, frequently forming small plates; branchlets pinkish brown, turning green-maroon, and later gray-maroon, 2 proximal internodes densely pubescent, distal internodes sparsely appressed-hairy; lenticels pale lenticular spots on new growth, uniformly scattered, especially on the distal internodes, protruding and extruding corky tissue on 2d year branches; pith white in 1st year branches, brown in older branches. |
Leaves | at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent; distalmost leaves much bigger than those at 2 more proximal nodes; petiole 0–3.4 mm; blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy; secondary veins 3 per side, all arising from proximal 1/2. |
petiole 2–8 mm; blade lanceolate to ovate, 1.5–8 × 0.5–4 cm, base cuneate, apex acuminate to obtuse, abaxial surface pale green, adaxial surface green, turning maroon in full sun, both surfaces with hairs appressed, white to ferruginous; secondary veins 3–4(–5 on some sucker shoots) per side, evenly spaced. |
Inflorescences | 20–40-flowered; peduncle 13–30 mm; primary branches 0–2 mm; bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate. |
usually elongate, sometimes convex or pyramidal, 2.5–5 cm diam., peduncle 20–40 mm; branches and pedicels yellow-green, turning bright red. |
Pedicels | 0.4–1.6 mm, sparsely appressed-hairy or glabrous. |
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Flowers | hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy; sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular; petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm; nectary dark purple or black. |
hypanthium appressed-hairy; sepals 0.2–0.5 mm; petals white, 2.3–3 mm. |
Drupes | 10–20 per inflorescence, red, globose, 6–8 mm; stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed. |
usually white, occasionally pale blue, oblate-ellipsoid, laterally compressed, 4–8 mm diam.; stone oblate-ellipsoid, 3–6.5 mm diam., smooth, apex rounded to slightly pointed. |
2n | = 44. |
= 22. |
Cornus unalaschkensis |
Cornus racemosa |
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Phenology | Flowering May–Aug; fruiting Aug–Oct. | Flowering May–Jul; fruiting Aug–Oct. |
Habitat | Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. | Fields, meadows, roadsides, fencerows, swamp margins. |
Elevation | 0–3000 m. (0–9800 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AK; CA; ID; OR; WA; AB; BC; YT
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CT; DC; IA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NE; NH; NJ; NY; OH; PA; RI; SD; TX; VA; VT; WI; WV; MB; ON; QC
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Discussion | Cornus racemosa, much like C. foemina and C. obliqua, has a coronulate or minutely papillate abaxial leaf surface, visible under high magnification. Cornus gracilis may be a hybrid between C. racemosa and C. foemina. Hybrids between C. obliqua and C. racemosa and C. rugosa are discussed under those species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 448. | FNA vol. 12, p. 456. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis | C. albida, C. foemina subsp. racemosa, C. gracilis, C. paniculata, C. paniculata var. albida, C. paniculata var. radiata, Swida candidissima, S. racemosa |
Name authority | Ledebour: Fl. Ross. 2: 378. (1844) | Lamarck: in J. Lamarck et al., Encycl. 2: 116. (1786) |
Web links |