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alaskan bunchberry, western bunchberry, western cordilleran bunchberry

mountain dogwood, Nuttall's dogwood, Pacific dogwood, western flowering dogwood

Habit Shrubs or trees to 20 m, flowering at 2 m.
Stems

erect, green, 6–22 cm, appressed-hairy;

nodes 4–6, internodes progressively longer distally;

branches only at distalmost node, much shorter than distal internodes so stems appear unbranched.

clustered, bark forming square to rectangular plates 0.5–1 cm wide;

branchlets green, maroon, or dark red, appressed-hairy;

lenticels pale round or lenticular spots, with dark central pore.

Leaves

at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent;

distalmost leaves much bigger than those at 2 more proximal nodes;

petiole 0–3.4 mm;

blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy;

secondary veins 3 per side, all arising from proximal 1/2.

petiole 5–17 mm;

blade elliptic, ovate, or obovate, 6–15 × 3–9.5 cm, base broadly cuneate, apex short acuminate, abaxial surface pale green, appressed-hairy, tufts of erect hairs present in axils of secondary veins, adaxial surface yellow green, appressed-hairy;

secondary veins 4–6 per side, most arising from proximal 1/2.

Inflorescences

20–40-flowered;

peduncle 13–30 mm;

primary branches 0–2 mm;

bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate.

convex, 1.5–3 cm diam., 40–75-flowered, subtended by pair of reduced leaves and pair of cataphylls;

peduncle 40–60 mm;

petaloid bracts 4–6, surrounding but not enclosing inflorescence through winter, white or tinged with pink, rhombic to obovate, 2.5–7 × 1.7–6 cm, apex short acuminate.

Pedicels

0.4–1.6 mm, sparsely appressed-hairy or glabrous.

Flowers

hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy;

sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular;

petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm;

nectary dark purple or black.

hypanthium appressed-hairy;

sepals 1–1.5 mm;

petals cream or yellow-green, rarely purple distally, 3–4 mm.

Drupes

10–20 per inflorescence, red, globose, 6–8 mm;

stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed.

red, closely appressed to each other, angular in cross section, 10–17 × 5–7.5 mm;

stone ellipsoid, 7–9 × 4–5 mm, smooth.

2n

= 44.

= 22.

Cornus unalaschkensis

Cornus nuttallii

Phenology Flowering May–Aug; fruiting Aug–Oct. Flowering Apr–Jul, sometimes Sep–Oct; fruiting Aug–Oct.
Habitat Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. Dry to moist coastal coniferous and montane forests.
Elevation 0–3000 m. (0–9800 ft.) 0–2400(–3000) m. (0–7900(–9800) ft.)
Distribution
from FNA
AK; CA; ID; OR; WA; AB; BC; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Source FNA vol. 12, p. 448. FNA vol. 12, p. 445.
Parent taxa Cornaceae > Cornus > subg. Arctocrania Cornaceae > Cornus > subg. Cynoxylon
Sibling taxa
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica, C. unalaschkensis
Synonyms Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis Benthamidia nuttallii, Cynoxylon nuttallii
Name authority Ledebour: Fl. Ross. 2: 378. (1844) Audubon: Ornithol. Biogr. 4: 482. (1838) — (as nuttalli)
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