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alaskan bunchberry, western bunchberry, western cordilleran bunchberry

Japanese dogwood, kousa, kousa dogwood

Habit Trees to 10 m, flowering at 2 m. Stems solitary or clustered, bark mottled gray and tan, of thin exfoliating plates; branches gray-maroon with longitudinal fissures, bark flaky; branchlets gray, densely appressed-hairy; lenticels pale, round or lenticular spots.
Stems

erect, green, 6–22 cm, appressed-hairy;

nodes 4–6, internodes progressively longer distally;

branches only at distalmost node, much shorter than distal internodes so stems appear unbranched.

Leaves

at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent;

distalmost leaves much bigger than those at 2 more proximal nodes;

petiole 0–3.4 mm;

blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy;

secondary veins 3 per side, all arising from proximal 1/2.

petiole 5–12 mm;

blade elliptic, ovate, or widely ovate, 5–8 × 2–5 cm, base cuneate, apex long acuminate, abaxial surface pale green, appressed-hairy, tufts of erect hairs in axils of secondary veins, adaxial surface dark green, appressed-hairy;

secondary veins 4–6 per side, evenly spaced, vein also present on leaf margin.

Inflorescences

20–40-flowered;

peduncle 13–30 mm;

primary branches 0–2 mm;

bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate.

subglobose, 0.9–1.4 cm diam., 40–75-flowered;

peduncle 40–60 mm;

petaloid bracts white to slightly yellow, narrowly ovate, rhombic, or trullate, 4–7 × 3–6 cm, base cuneate, apex acuminate.

Pedicels

0.4–1.6 mm, sparsely appressed-hairy or glabrous.

Flowers

hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy;

sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular;

petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm;

nectary dark purple or black.

hypanthium appressed-hairy;

sepals 1–1.5 mm;

petals cream or yellow-green, 3–4 mm.

Drupes

10–20 per inflorescence, red, globose, 6–8 mm;

stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed.

Syncarps

red, subglobose, 20–27 mm diam.;

stones ellipsoid, 7–9 × 4–5 mm, smooth.

2n

= 44.

Cornus unalaschkensis

Cornus kousa

Phenology Flowering May–Aug; fruiting Aug–Oct. Flowering Apr–Jul; fruiting Aug–Oct.
Habitat Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. Disturbed areas, lawns.
Elevation 0–3000 m. (0–9800 ft.) 0–1000 m. (0–3300 ft.)
Distribution
from FNA
AK; CA; ID; OR; WA; AB; BC; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
NC; NY; Asia (China, Japan, Korea) [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Cornus kousa is frequently planted in the flora area as an ornamental and for its edible fruit. Though not well established, it has been noted to spread from cultivation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 448. FNA vol. 12, p. 445.
Parent taxa Cornaceae > Cornus > subg. Arctocrania Cornaceae > Cornus > subg. Syncarpea
Sibling taxa
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica, C. unalaschkensis
Synonyms Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis Benthamia japonica, Benthamidia japonica, Dendrobenthamia japonica
Name authority Ledebour: Fl. Ross. 2: 378. (1844) Bürger ex Hance: J. Linn. Soc., Bot. 13: 105. (1872)
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