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alaskan bunchberry, western bunchberry, western cordilleran bunchberry

brown dogwood, smooth dogwood

Habit Shrubs, to 3 m, flowering at 1 m; rhizomes absent.

erect, green, 6–22 cm, appressed-hairy;

nodes 4–6, internodes progressively longer distally;

branches only at distalmost node, much shorter than distal internodes so stems appear unbranched.

clustered, branches at colony margin decumbent or trailing and rooting at the nodes, forming large thickets;

bark gray, flaky, not corky, appearing braided, splitting longitudinally;

branchlets brown, pink, or maroon, appressed-hairy;

lenticels inconspicuous, swelling but seldom protruding or extruding tissue on 2d year branches, periderm swelling around them and often over large contiguous areas;

pith brown.


at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent;

distalmost leaves much bigger than those at 2 more proximal nodes;

petiole 0–3.4 mm;

blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy;

secondary veins 3 per side, all arising from proximal 1/2.

petiole 3–7 mm;

blade lanceolate, elliptic, or obovate, 3–5 × 1.5–2.5 cm, base cuneate, apex abruptly acuminate or rounded, abaxial surface pale green, adaxial surface gray-green, both surfaces with hairs appressed, sparse;

secondary veins 3–4(–5) per side, evenly spaced, basal vein arising 5–10 mm from blade base.



peduncle 13–30 mm;

primary branches 0–2 mm;

bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate.

flat-topped or slightly convex, 2.5–4.5 cm diam., peduncle 10–20 mm;

branches and pedicels green or maroon.


0.4–1.6 mm, sparsely appressed-hairy or glabrous.


hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy;

sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular;

petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm;

nectary dark purple or black.

hypanthium densely appressed-hairy;

sepals 0.4–0.8 mm;

petals cream, 3–5 mm.


10–20 per inflorescence, red, globose, 6–8 mm;

stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed.

blue, bleaching white in direct sun, globose or subglobose, 6–9 mm diam.;

stone globose to subglobose, 4–6 mm diam., smooth, apex rounded.


= 44.

= 22.

Cornus unalaschkensis

Cornus glabrata

Phenology Flowering May–Aug; fruiting Aug–Oct. Flowering May–Jun; fruiting Aug–Oct.
Habitat Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. Stream banks, roadsides, fields, meadows.
Elevation 0–3000 m. (0–9800 ft.) 50–1500 m. (200–4900 ft.)
from FNA
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[BONAP county map]
from FNA
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[BONAP county map]

Cornus glabrata, which ranges from southern Oregon through most of transmontane California, grows in two habits. Along streams and in moist areas, the branches are trailing, often vinelike, and rooting at the nodes. H. McMinn (1939) reported this growth habit for cultivated plants. A second form is found in drier roadsides and fields, where the stems are erect or grow horizontally, rooting at the nodes and forming dense thickets. This growth pattern is accompanied by smaller, thicker leaves and erect branches. There is no doubt this is a single species, because the authors have observed a single clone on a stream bank, with half the clone growing erect, with small leaves, and half trailing into the stream bed, with large leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 448. FNA vol. 12, p. 457.
Parent taxa Cornaceae > Cornus > subg. Arctocrania Cornaceae > Cornus > subg. Thelycrania
Sibling taxa
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica, C. unalaschkensis
Synonyms Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis C. costulata, Swida catalinensis
Name authority Ledebour: Fl. Ross. 2: 378. (1844) Bentham: Bot. Voy. Sulphur, 18. (1844)
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