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alaskan bunchberry, western bunchberry, western cordilleran bunchberry

stiff dogwood, swamp dogwood

Habit Shrubs, to 8 m, flowering at 1.5 m; rhizomes absent.
Stems

erect, green, 6–22 cm, appressed-hairy;

nodes 4–6, internodes progressively longer distally;

branches only at distalmost node, much shorter than distal internodes so stems appear unbranched.

clustered;

bark gray-brown, becoming gray-black, corky, appearing braided, splitting longitudinally, checkered;

branchlets deep red, often pale green abaxially, or completely green or bronze if shaded, 2 proximal internodes densely pubescent, distal internodes sparsely appressed-hairy;

lenticels pale circular spots on new growth, usually more common on distal portion of internodes, often overlapping, localized to form longitudinal bands, splitting periderm but not protruding or extruding tissue on 2d year branches, periderm swelling around them and usually over large contiguous areas;

pith white in 1st year branches, tan in older branches.

Leaves

at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent;

distalmost leaves much bigger than those at 2 more proximal nodes;

petiole 0–3.4 mm;

blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy;

secondary veins 3 per side, all arising from proximal 1/2.

petiole 5–16 mm;

blade lanceolate, elliptic, or oblanceolate, 3.5–11 × 1–6 cm, base cuneate to rounded, apex abruptly acuminate to an obtuse tip, abaxial surface pale green, adaxial surface dark green, midvein turning red or maroon, both surfaces with hairs appressed, sparse, glabrate by late summer;

secondary veins 3–4 per side, evenly spaced, basal vein arising 1–2 mm from blade base.

Inflorescences

20–40-flowered;

peduncle 13–30 mm;

primary branches 0–2 mm;

bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate.

flat-topped, convex, or pyramidal, 2–8 cm diam., peduncle 15–45 mm;

branches and pedicels greenish yellow, turning red in fruit.

Pedicels

0.4–1.6 mm, sparsely appressed-hairy or glabrous.

Flowers

hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy;

sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular;

petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm;

nectary dark purple or black.

hypanthium densely appressed-hairy;

sepals 0.4–1 mm;

petals cream, 2.6–3.8 mm.

Drupes

10–20 per inflorescence, red, globose, 6–8 mm;

stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed.

pale blue, violet plumbeous, or blue violet, often turning whitish blue to white, globose or oblate-ellipsoid, 5–6 × 6–10 mm;

stone globose to oblong, 3–3.7 × 3.7–5 mm, slightly ribbed, apex rounded.

2n

= 44.

Cornus unalaschkensis

Cornus foemina

Phenology Flowering May–Aug; fruiting Aug–Oct. Flowering Mar–Jun; fruiting Aug–Oct.
Habitat Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. Marshes, swamps, river and stream banks, pocosin margins, interdune swales, wet ditches.
Elevation 0–3000 m. (0–9800 ft.) 0–1500 m. (0–4900 ft.)
Distribution
from FNA
AK; CA; ID; OR; WA; AB; BC; YT
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[BONAP county map]
from FNA
AL; AR; DE; FL; GA; IL; IN; KY; LA; MD; MO; MS; NC; NJ; OK; SC; TN; TX; VA
[WildflowerSearch map]
[BONAP county map]
Discussion

Cornus foemina has a coronulate or minutely papillate abaxial leaf surface, visible under high magnification, like that of C. obliqua and C. racemosa.

Cornus cyanocarpus J. F. Gmelin 1791, a parahomonym (thus illegitimate) of C. cyanocarpa Moench 1785, pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 448. FNA vol. 12, p. 456.
Parent taxa Cornaceae > Cornus > subg. Arctocrania Cornaceae > Cornus > subg. Thelycrania
Sibling taxa
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica, C. unalaschkensis
Synonyms Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis C. stricta, Swida foemina, S. stricta
Name authority Ledebour: Fl. Ross. 2: 378. (1844) Miller: Gard. Dict. ed. 8, Cornus no. 4. (1768)
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