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alaskan bunchberry, western bunchberry, western cordilleran bunchberry

flowering dogwood

Habit Trees to 20 m, flowering at 2 m.
Stems

erect, green, 6–22 cm, appressed-hairy;

nodes 4–6, internodes progressively longer distally;

branches only at distalmost node, much shorter than distal internodes so stems appear unbranched.

clustered, occasionally decumbent and rooting at nodes, bark corky, forming rectangular plates 0.5–1 cm wide;

branchlets green, maroon, or red, appressed-hairy;

lenticels maroon swellings.

Leaves

at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent;

distalmost leaves much bigger than those at 2 more proximal nodes;

petiole 0–3.4 mm;

blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy;

secondary veins 3 per side, all arising from proximal 1/2.

petiole 3–20 mm;

blade ovate, elliptic, or obovate, 5–12 × 2–7 cm, base cuneate to rounded, apex abruptly acuminate, abaxial surface whitish, appressed-hairy, tufts of erect hairs present in axils of secondary veins, adaxial surface dark green, appressed-hairy;

secondary veins 5–7 per side, most arising from proximal 1/2.

Inflorescences

20–40-flowered;

peduncle 13–30 mm;

primary branches 0–2 mm;

bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate.

flat-topped, 1–2 cm diam., 15–30-flowered, subtended by 2 pairs of cataphylls;

peduncle 10–20 mm;

petaloid bracts 4, surrounding and enclosing inflorescence through winter, white or tinged with red and with brown or white callous at apex, obovate to obcordate, 2–6 × 1–4.5 cm, apex rounded or emarginate.

Pedicels

0.4–1.6 mm, sparsely appressed-hairy or glabrous.

Flowers

hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy;

sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular;

petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm;

nectary dark purple or black.

hypanthium appressed-hairy;

sepals 0.5–0.8 mm;

petals cream or yellow-green, 3–3.5 mm.

Drupes

10–20 per inflorescence, red, globose, 6–8 mm;

stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed.

usually red, rarely yellow, drying black, spreading from each other, round in cross section, 13–18 × 6–9 mm;

stone ellipsoid, 10–12 × 4–7 mm, smooth.

2n

= 44.

= 44.

Cornus unalaschkensis

Cornus florida

Phenology Flowering May–Aug; fruiting Aug–Oct. Flowering Mar–Jun; fruiting Aug–Oct.
Habitat Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. Deciduous, mixed, and pine forests.
Elevation 0–3000 m. (0–9800 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AK; CA; ID; OR; WA; AB; BC; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WV; ON; e Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Mexican populations of Cornus florida sometimes have been treated as C. urbinia Rose or C. florida subsp. urbinia (Rose) Rickett, distinguished primarily by bract size, shape, and apical cohesion following expansion. North American and Mexican populations overlap in these characters and are treated here as a single taxon.

Cornus florida, the state tree of Missouri and Virginia and the state flower of North Carolina, is an understory tree that can form spectacular displays when flowering. Cultivars with pink to red bracts are often planted as ornamentals. Dogwood anthracnose is causing serious declines in C. florida throughout its range.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 448. FNA vol. 12, p. 446.
Parent taxa Cornaceae > Cornus > subg. Arctocrania Cornaceae > Cornus > subg. Cynoxylon
Sibling taxa
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica, C. unalaschkensis
Synonyms Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis Benthamidia florida, Cynoxylon floridum
Name authority Ledebour: Fl. Ross. 2: 378. (1844) Linnaeus: Sp. Pl. 1: 117. (1753)
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