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alaskan bunchberry, western bunchberry, western cordilleran bunchberry

rough-leaf dogwood

Habit Shrubs or trees, to 6 m, flowering at 1.5 m; rhizomes present.
Stems

erect, green, 6–22 cm, appressed-hairy;

nodes 4–6, internodes progressively longer distally;

branches only at distalmost node, much shorter than distal internodes so stems appear unbranched.

solitary, 1–6 dm apart;

bark pinkish gray, appearing braided, becoming corky, checkered with square or rectangular plates 2–5 mm wide;

branchlets yellow-green abaxially, pink-maroon adaxially, densely erect-hairy when young;

lenticels inconspicuous on new growth, protruding and splitting longitudinally on 2d year branches;

pith brown, white, or tan.

Leaves

at proximal 2–4 nodes nonchlorophyllous, opposite, ± scalelike, caducous (rarely chlorophyllous at 3d node from apex but much smaller than more distal leaves), at 2d node from apex nonchlorophyllous proximally, chlorophyllous distally, opposite, well developed, persistent, at distalmost node chlorophyllous, appearing to be in whorl of 6, well developed, persistent;

distalmost leaves much bigger than those at 2 more proximal nodes;

petiole 0–3.4 mm;

blade ovate to elliptic, 3.5–8 × 0.9–4 cm, apex acute or short acuminate, abaxial surface pale green, hairs sparsely appressed-hairy, adaxial surface green, appressed-hairy;

secondary veins 3 per side, all arising from proximal 1/2.

petiole 8–25 mm;

blade lanceolate to ovate, 2–12 × 1.2–7.7 cm, base cuneate, truncate, or cordate, apex abruptly acuminate, abaxial surface pale green, hairs curved upward, dense, adaxial surface gray-green, hairs curved upward or appressed;

secondary veins 3–4(–5) per side, most arising from proximal 1/2.

Inflorescences

20–40-flowered;

peduncle 13–30 mm;

primary branches 0–2 mm;

bracts greenish white or white, often red-tipped, unequal, 2 ovate, 21–30 × 12–13 mm, 2 suborbiculate, 17–1.9 × 13–16 mm, apex acuminate.

flat-topped or convex, 3–8 cm diam., peduncle 20–40 mm;

branches and pedicels pink or red, fading yellow when dried, central part of pedicels white after fruit falls.

Pedicels

0.4–1.6 mm, sparsely appressed-hairy or glabrous.

Flowers

hypanthium cream to mottled purple, 1.2–2 mm, densely appressed-hairy;

sepals mottled purple and cream, 0.1–0.4 mm, apex rounded or acute, thick, sparsely hairy on margin, densely glandular;

petals cream proximally, purple distally, 1.5–1.8 mm, apical awn 0.4–0.6 mm;

nectary dark purple or black.

hypanthium densely appressed-hairy;

sepals 0.4–1.3 mm;

petals white, 2.3–3.2 mm.

Drupes

10–20 per inflorescence, red, globose, 6–8 mm;

stone globose or subglobose, 2.7–3.4 × 2.1–3.4 mm, longitudinally grooved, apex slightly pointed.

white, globose or subglobose, 4–7.5 mm diam.;

stone subglobose, 3–6 mm diam., smooth or slightly grooved, apex rounded.

2n

= 44.

= 22.

Cornus unalaschkensis

Cornus drummondii

Phenology Flowering May–Aug; fruiting Aug–Oct. Flowering Apr–Jul; fruiting Aug–Oct.
Habitat Maritime copse or heath, maritime coniferous forests and bog woodlands, moist broadleaf or coniferous forests. Limestone barrens, limestone outcrops, dry woodlands, rocky stream banks, prairies, old fields, roadsides, meadows, swamp margins.
Elevation 0–3000 m. (0–9800 ft.) 0–1500 m. (0–4900 ft.)
Distribution
from FNA
AK; CA; ID; OR; WA; AB; BC; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; GA; IA; IL; IN; KS; KY; LA; MI; MO; MS; NE; NY; OH; OK; PA; SD; TN; TX; WI; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Several workers considered the erect hairs on the adaxial leaf surface of Cornus drummondii and C. asperifolia to indicate relatedness, and treated these two species as one (J. Torrey and A. Gray 1838–1843; Gray 1856; J. M. Coulter and W. H. Evans 1890; W. Wangerin 1910). However, A. Wood (1861), following W. T. Feay, linked C. asperifolia with C. foemina, and J. S. Wilson (1964) also segregated the southern coastal plain rough-leaved dogwood from the continental rough-leaved dogwood as C. foemina subsp. microcarpa (Nash) J. S. Wilson and C. drummondii, respectively.

J. K. Small described Cornus priceae from Bowling Green, Kentucky, with small drupes and stones. However, extensive collections from the type locality and examination of the type material revealed that the drupe and stone sizes are within the normal range of variation of C. drummondii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 448. FNA vol. 12, p. 455.
Parent taxa Cornaceae > Cornus > subg. Arctocrania Cornaceae > Cornus > subg. Thelycrania
Sibling taxa
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. drummondii, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica
C. alternifolia, C. amomum, C. asperifolia, C. canadensis, C. florida, C. foemina, C. glabrata, C. kousa, C. mas, C. nuttallii, C. obliqua, C. occidentalis, C. racemosa, C. rugosa, C. sanguinea, C. sericea, C. sessilis, C. suecica, C. unalaschkensis
Synonyms Arctocrania unalaschkensis, Chamaepericlymenum unalaschkense, Cornella unalaschkensis, Swida unalaschkensis C. asperifolia var. drummondii, C. priceae, Swida drummondii, S. priceae
Name authority Ledebour: Fl. Ross. 2: 378. (1844) C. A. Meyer: Cornus-Arten, 20. (1845) — (as drummondi)
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