FNA: Corispermum pacificum vs. Corispermum navicula

	Corispermum pacificum	Corispermum navicula
	common bugseed, Pacific bug-seed	boat-shape bugseed, crescent bugseed
Habit	Plants usually branched from base, (5–)15–40 cm, glabrous or sparsely covered with dendroid hairs (especially when young).	Plants branched from the base or nearly so, 5–15(–25) cm, sparsely covered with dendroid or stellate hairs, or almost glabrous.
Leaf	blades narrowly lanceolate, linear-lanceolate, or linear, flat or nearly so, 2–5(–7) × 0.2–0.6 cm.	blades linear-lanceolate, linear, occasionally narrowly lanceolate, usually plane, (1.5–)2–4 × 0.1–0.5 cm.
Bracts	ovate-lanceolate, narrowly ovate-lanceolate, almost ovate, or lanceolate, (1–)1.5–2.5 × 0.3–0.7(–0.9) cm.	ovate or ovate-lanceolate (occasionally proximal ones leaflike, narrowly ovate-lanceolate or lanceolate), 0.5–2 × 0.2–0.6 cm.
Inflorescences	usually compact, rather dense, or sometimes ± lax, condensed only near apex, ovoid, oblong-ovate, obovate, broadly linear, or occasionally clavate.	compact and dense, ovoid, ovate or oblong-obovate.
Perianth	segment 1, sometimes absent in distal flowers.	segment 1.
Fruits	usually black (rarely deep olive green, especially when immature), sharply contrasting with greenish semitransparent wings, without spots and warts, slightly convex abaxially, flat or slightly concave adaxially, orbiculate-obovate to almost orbiculate, broadest near middle (or occasionally slightly beyond), 3–4 × 2.7–3.8 mm, shiny; wing translucent, thin, (0.2–)0.3–0.6 mm wide, margins slightly undulate or indistinctly erose-denticulate, apex rounded or occasionally indistinctly notched.	brown, dark brown, or deep olive green, usually with numerous reddish brown spots and whitish warts, strongly convex abaxially, usually strongly concave adaxially, elongate-obovate or obovate-elliptic, broadest beyond middle, (4.2–)4.5–5(–5.2) × 2.5–3 mm; wing not translucent or translucent only at margin, thick, 0.1–0.2(–0.3) mm wide (occasionally nearly absent), margins entire or irregularly erose, usually involute toward adaxial face of fruit, apex rostrate, triangular (wing long-adnate to style bases).

	Corispermum pacificum	Corispermum navicula
Phenology	Flowering late summer–fall.	Flowering late summer–fall.
Habitat	Sandy shores, dunes	Sand dunes, probably also sandy and gravely shores
Elevation	0-500 m (0-1600 ft)	2500 m (8200 ft)
Distribution	ID; OR; WA [WildflowerSearch map] [BONAP county map]	CO [BONAP county map]
Discussion	Of conservation concern. Corispermum pacificum seems to be closely related to Siberian C. crassifolium Turczaninov and C. maynense Ignatov. The latter species occurs in the northeastern Russian Far East and may be expected to occur in Alaska. Corispermum pacificum differs from C. maynense by its usually more robust habit, and its wing rounded (rarely rounded-truncate or indistinctly emarginate, but not triangular) at apex. From C. crassifolium it may be distinguished by the constant presence of perianth segments, and more flattened black mature fruits. Corispermum pacificum probably also occurs in adjacent regions of British Columbia. Corispermum pacificum is placed in subsect. Crassifolia (S. L. Mosyakin 1997). This subsection seems to be of Siberian origin, with its central species, C. crassifolium, being closest to the hypothetic ancestral taxon. The presence of perianth segments in C. pacificum may be explained by ancient hybridization with representatives of subsect. Pallasiana. Reproductive isolation between the sympatric species of Corispermum may be achieved by different flowering periods. Occasional hybrids between C. pacificum and C. villosum are similar in their habit to C. pacificum in having usually broad leaves and rather dense inflorescences but they have mostly aborted fruits suggesting that C. pacificum and C. villosum are taxonomically distant species. (Discussion copyrighted by Flora of North America; reprinted with permission.)	Corispermum navicula is very similar in its fruit morphology to the Siberian species C. bardunovii M. Popov ex M. Lomonosova (M. N. Lomonosova 1992). Probably, the two taxa represent results of parallel evolution (or parallel variability?) within North American and Asian representatives of the same species aggregate. The most distinctive character of both C. navicula and C. bardunovii, an elongated fruit body with almost parallel margins in the middle portion and distinctly triangular apex, shows a transition toward representatives of Corispermum sect. Declinata Mosyakin. Additional study of C. navicula would help clarify its relationships with other species. Some specimens from Oklahoma may also belong to C. navicula. (Discussion copyrighted by Flora of North America; reprinted with permission.)
Source	FNA vol. 4, p. 319.	FNA vol. 4, p. 317.
Parent taxa	Chenopodiaceae > Corispermum	Chenopodiaceae > Corispermum
Sibling taxa	C. americanum, C. hookeri, C. hyssopifolium, C. navicula, C. nitidum, C. ochotense, C. pallasii, C. pallidum, C. villosum, C. welshii	C. americanum, C. hookeri, C. hyssopifolium, C. nitidum, C. ochotense, C. pacificum, C. pallasii, C. pallidum, C. villosum, C. welshii
Name authority	Mosyakin: Novon 5: 345, fig. 1A. (1995)	Mosyakin: Novon 5: 349. (1995)
Web links	Local floras: BC, OR, WA Local Web sites: Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Corispermum navicula

common bugseed, Pacific bug-seed

boat-shape bugseed, crescent bugseed

Habit

Plants usually branched from base, (5–)15–40 cm, glabrous or sparsely covered with dendroid hairs (especially when young).

Plants branched from the base or nearly so, 5–15(–25) cm, sparsely covered with dendroid or stellate hairs, or almost glabrous.

Leaf

blades narrowly lanceolate, linear-lanceolate, or linear, flat or nearly so, 2–5(–7) × 0.2–0.6 cm.

blades linear-lanceolate, linear, occasionally narrowly lanceolate, usually plane, (1.5–)2–4 × 0.1–0.5 cm.

Bracts

ovate-lanceolate, narrowly ovate-lanceolate, almost ovate, or lanceolate, (1–)1.5–2.5 × 0.3–0.7(–0.9) cm.

ovate or ovate-lanceolate (occasionally proximal ones leaflike, narrowly ovate-lanceolate or lanceolate), 0.5–2 × 0.2–0.6 cm.

Inflorescences

usually compact, rather dense, or sometimes ± lax, condensed only near apex, ovoid, oblong-ovate, obovate, broadly linear, or occasionally clavate.

compact and dense, ovoid, ovate or oblong-obovate.

Perianth

segment 1, sometimes absent in distal flowers.

segment 1.

Fruits

usually black (rarely deep olive green, especially when immature), sharply contrasting with greenish semitransparent wings, without spots and warts, slightly convex abaxially, flat or slightly concave adaxially, orbiculate-obovate to almost orbiculate, broadest near middle (or occasionally slightly beyond), 3–4 × 2.7–3.8 mm, shiny;

wing translucent, thin, (0.2–)0.3–0.6 mm wide, margins slightly undulate or indistinctly erose-denticulate, apex rounded or occasionally indistinctly notched.

brown, dark brown, or deep olive green, usually with numerous reddish brown spots and whitish warts, strongly convex abaxially, usually strongly concave adaxially, elongate-obovate or obovate-elliptic, broadest beyond middle, (4.2–)4.5–5(–5.2) × 2.5–3 mm;

wing not translucent or translucent only at margin, thick, 0.1–0.2(–0.3) mm wide (occasionally nearly absent), margins entire or irregularly erose, usually involute toward adaxial face of fruit, apex rostrate, triangular (wing long-adnate to style bases).

Corispermum pacificum

Corispermum navicula

Phenology

Flowering late summer–fall.

Habitat

Sandy shores, dunes

Sand dunes, probably also sandy and gravely shores

Elevation

0-500 m (0-1600 ft)

2500 m (8200 ft)

Distribution

ID; OR; WA

[WildflowerSearch map]

[BONAP county map]

Discussion

Of conservation concern.

Corispermum pacificum seems to be closely related to Siberian C. crassifolium Turczaninov and C. maynense Ignatov. The latter species occurs in the northeastern Russian Far East and may be expected to occur in Alaska. Corispermum pacificum differs from C. maynense by its usually more robust habit, and its wing rounded (rarely rounded-truncate or indistinctly emarginate, but not triangular) at apex. From C. crassifolium it may be distinguished by the constant presence of perianth segments, and more flattened black mature fruits. Corispermum pacificum probably also occurs in adjacent regions of British Columbia. Corispermum pacificum is placed in subsect. Crassifolia (S. L. Mosyakin 1997). This subsection seems to be of Siberian origin, with its central species, C. crassifolium, being closest to the hypothetic ancestral taxon. The presence of perianth segments in C. pacificum may be explained by ancient hybridization with representatives of subsect. Pallasiana.

Reproductive isolation between the sympatric species of Corispermum may be achieved by different flowering periods. Occasional hybrids between C. pacificum and C. villosum are similar in their habit to C. pacificum in having usually broad leaves and rather dense inflorescences but they have mostly aborted fruits suggesting that C. pacificum and C. villosum are taxonomically distant species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Corispermum navicula is very similar in its fruit morphology to the Siberian species C. bardunovii M. Popov ex M. Lomonosova (M. N. Lomonosova 1992). Probably, the two taxa represent results of parallel evolution (or parallel variability?) within North American and Asian representatives of the same species aggregate. The most distinctive character of both C. navicula and C. bardunovii, an elongated fruit body with almost parallel margins in the middle portion and distinctly triangular apex, shows a transition toward representatives of Corispermum sect. Declinata Mosyakin. Additional study of C. navicula would help clarify its relationships with other species. Some specimens from Oklahoma may also belong to C. navicula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source

FNA vol. 4, p. 319.

FNA vol. 4, p. 317.

Parent taxa

Chenopodiaceae > Corispermum

Sibling taxa

C. americanum, C. hookeri, C. hyssopifolium, C. navicula, C. nitidum, C. ochotense, C. pallasii, C. pallidum, C. villosum, C. welshii

C. americanum, C. hookeri, C. hyssopifolium, C. nitidum, C. ochotense, C. pacificum, C. pallasii, C. pallidum, C. villosum, C. welshii

Name authority

Mosyakin: Novon 5: 345, fig. 1A. (1995)

Mosyakin: Novon 5: 349. (1995)

Web links

Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).

Choose a data source: Flora of North America, OregonFlora, UW Burke Herbarium

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Corispermum pacificum

Corispermum navicula

Corispermum pacificum

Corispermum navicula