Corispermum pacificum |
Corispermum hyssopifolium |
|
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common bugseed, Pacific bug-seed |
bugseed, hyssop-leaf bugseed |
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Habit | Plants usually branched from base, (5–)15–40 cm, glabrous or sparsely covered with dendroid hairs (especially when young). | Plants branched from base (rarely slightly above base), 10–35(–55) cm, densely to sparsely covered with dendroid or stellate hairs, becoming glabrous. |
Leaf | blades narrowly lanceolate, linear-lanceolate, or linear, flat or nearly so, 2–5(–7) × 0.2–0.6 cm. |
blades linear-lanceolate or linear; usually plane,1.5–3.5(–4) × 0.2–0.4(–0.5) cm. |
Bracts | ovate-lanceolate, narrowly ovate-lanceolate, almost ovate, or lanceolate, (1–)1.5–2.5 × 0.3–0.7(–0.9) cm. |
ovate or ovate-lanceolate (rarely narrowly ovate-lanceolate to ± lanceolate), 0.5–1.5(–2) × 0.3–0.7 cm. |
Inflorescences | usually compact, rather dense, or sometimes ± lax, condensed only near apex, ovoid, oblong-ovate, obovate, broadly linear, or occasionally clavate. |
compact, usually not strongly condensed at apex, occasionally interrupted near base, linear, oblong-linear, or indistinctly clavate-linear. |
Perianth | segment 1, sometimes absent in distal flowers. |
segments 1(–3). |
Fruits | usually black (rarely deep olive green, especially when immature), sharply contrasting with greenish semitransparent wings, without spots and warts, slightly convex abaxially, flat or slightly concave adaxially, orbiculate-obovate to almost orbiculate, broadest near middle (or occasionally slightly beyond), 3–4 × 2.7–3.8 mm, shiny; wing translucent, thin, (0.2–)0.3–0.6 mm wide, margins slightly undulate or indistinctly erose-denticulate, apex rounded or occasionally indistinctly notched. |
brown, dark brown, or deep olive green, usually without spots and warts, broadly elliptic, prominently convex abaxially, usually plane or slightly concave adaxially, obovate-elliptic, or ± orbiculate, broadest near middle (rarely slightly beyond middle), 2.2–3.2 (–3.5) × 1.7–2.8 mm shiny wing (when present) translucent at margins, 0.1(–0.15) mm wide, margins entire, apex rounded. |
Corispermum pacificum |
Corispermum hyssopifolium |
|
Phenology | Flowering late summer–fall. | Flowering summer–early fall. |
Habitat | Sandy shores, dunes | Sandy waste places, roadsides, shores |
Elevation | 0-500 m (0-1600 ft) | elevation not known |
Distribution |
ID; OR; WA
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CO; se Europe; w Asia [Introduced in North America] |
Discussion | Of conservation concern. Corispermum pacificum seems to be closely related to Siberian C. crassifolium Turczaninov and C. maynense Ignatov. The latter species occurs in the northeastern Russian Far East and may be expected to occur in Alaska. Corispermum pacificum differs from C. maynense by its usually more robust habit, and its wing rounded (rarely rounded-truncate or indistinctly emarginate, but not triangular) at apex. From C. crassifolium it may be distinguished by the constant presence of perianth segments, and more flattened black mature fruits. Corispermum pacificum probably also occurs in adjacent regions of British Columbia. Corispermum pacificum is placed in subsect. Crassifolia (S. L. Mosyakin 1997). This subsection seems to be of Siberian origin, with its central species, C. crassifolium, being closest to the hypothetic ancestral taxon. The presence of perianth segments in C. pacificum may be explained by ancient hybridization with representatives of subsect. Pallasiana. Reproductive isolation between the sympatric species of Corispermum may be achieved by different flowering periods. Occasional hybrids between C. pacificum and C. villosum are similar in their habit to C. pacificum in having usually broad leaves and rather dense inflorescences but they have mostly aborted fruits suggesting that C. pacificum and C. villosum are taxonomically distant species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In addition to the key characteristics, Corispermum hyssopifolium is distinguished by having style bases short, barely exposed over the edge of wing and fruit surfaces which are glabrous and shiny. The application of the Linnaean name Corispermum hyssopifolium is problematic. The standing lectotype (C. E. Jarvis et al. 1993; I. C. Hedge 1997) and traditional circumscription of C. hyssopifolium (as understood by M. M. Iljin 1936; P. Aellen 1961, 1964; and many others) are accepted here. No unquestionable specimens of Corispermum hyssopifolium sensu stricto are known from North America. A collection from Colorado and some collections from Missouri approach this species most closely (S. L. Mosyakin 1995). However, their fruit morphology suggests that they are either members of the native Asian and North American group centered around C. pallasii, or resulted from hybridization between C. hyssopifolium and native representatives of subsect. Pallasiana. The name Corispermum hyssopifolium has been misapplied to the majority of species of the genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 4, p. 319. | FNA vol. 4. |
Parent taxa | Chenopodiaceae > Corispermum | Chenopodiaceae > Corispermum |
Sibling taxa | ||
Name authority | Mosyakin: Novon 5: 345, fig. 1A. (1995) | Linnaeus: Sp. Pl. 1: 4. (1753) |
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