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alaskan bugseed, Okhotia n bugseed, Russian bugseed

Welsh's bugseed

Habit Plants branched from the base or nearly so, 5–15(–20) cm, sparsely covered with dendroid or stellate hairs, becoming glabrous.
Leaf

blades linear-oblanceolate (linear-spatulate) or linear, often broadest in distal 1/3, plane, 1–3.5 × 0.1–0.3 cm, abruptly contracted into mucronulate apex.

blades linear-lanceolate or linear, usually plane (or rarely slightly convolute or folded in dried plants), 1–6 × 0.2–0.5 cm.

Bracts

ovate-lanceolate, lanceolate, or linear-lanceolate (proximal bracts usually linear, leaflike, 4–7 times fruit length), (0.5–)1–1.5(–3) × (0.1–)0.2–0.5 cm.

ovate or ovate-lanceolate (rarely narrowly ovate-lanceolate), 1–3 × 0.3–0.8 cm.

Inflorescences

compact and dense, short-clavate or almost ovoid (if linear, then all bracts leaflike).

usually compact and dense, rarely ± lax, and condensed only at apex, ovoid, oblong-obovate, or oblong-clavate.

Perianth

segment 1.

segment 1.

Fruits

reddish brown, dark brown, deep olive green (then often tinged with red), or deep beet red, usually without dark spots or whitish warts, strongly convex abaxially, plane or slightly concave adaxially, obovate-elliptic or almost obovate, usually broadest near middle (rarely slightly beyond middle), (2.5–)2.8–4 × 1.8–2.7 mm;

wing translucent only at margins, thick, (0.1–)0.2–0.3 mm wide, margins entire, apex broadly triangular or almost rounded.

yellowish brown, light brown, or brown, usually with reddish brown spots and whitish warts, strongly to slightly convex abaxially, usually concave adaxially, obovate or orbiculate-obovate, usually broadest slightly beyond middle (occasionally almost near middle), (3.3–)3.7–4.6 × (2.7–)3–3.6 mm, slightly shiny or dull;

wing translucent, thin, (rarely translucent only at margin, thicker), (0.3–)0.4–0.6 mm wide, margins entire or irregularly minutely erose-denticulate, apex rounded, truncate, or indistinctly emarginate.

Plant

branched from base or nearly so, 10–35 cm, densely or sparsely covered with dendroid or stellate hairs (rarely with scattered papillae at margins of bracts).

Corispermum ochotense

Corispermum welshii

Phenology Flowering late summer–fall.
Habitat Sand dunes, sandy shores of creeks and rivers, dry valleys
Elevation 1300-2200 m (4300-7200 ft)
Distribution
from FNA
AK; NT; Eurasia
[BONAP county map]
from FNA
AZ; NM; UT; WY
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Corispermum ochotense is conspicuous in that it is usually red at maturity. No specimens of Corispermum have been seen from Yukon Territory, but the species might be expected there. One collection of immature plants from northern Saskatchewan (see S. L. Mosyakin 1995) probably also belongs to C. ochotense. Specimens of C. ochotense have been misidentified in North America and northeast Asia as C. hyssopifolium Linnaeus and C. sibiricum Iljin.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The names Corispermum hyssopifolium and C. villosum have been misapplied to C. welshii. Judging from its fruit morphology, C. welshii is closely related to C. americanum, especially to var. rydbergii. Forms intermediate between these taxa occasionally occur, especially in Utah. However, C. welshii differs from C. americanum in having shorter and thicker inflorescences and broader leaves and bracts. Forms similar to C. welshii occur also in other southwestern states, particularly in Colorado and western Texas. These plants are in need of additional study.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits (2.5-)2.8-3.2 × 1.8-2.2 mm
var. ochotense
1. Fruits 3.2-4 × 2.2-2.7 mm
var. alaskanum
Source FNA vol. 4. FNA vol. 4, p. 317.
Parent taxa Chenopodiaceae > Corispermum Chenopodiaceae > Corispermum
Sibling taxa
C. americanum, C. hookeri, C. hyssopifolium, C. navicula, C. nitidum, C. pacificum, C. pallasii, C. pallidum, C. villosum, C. welshii
C. americanum, C. hookeri, C. hyssopifolium, C. navicula, C. nitidum, C. ochotense, C. pacificum, C. pallasii, C. pallidum, C. villosum
Subordinate taxa
C. ochotense var. alaskanum, C. ochotense var. ochotense
Name authority Ignatov: Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 91(3): 113, fig. 2ß. (1986) Mosyakin: Novon 5: 348, fig. 1E. (1995)
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