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leafstem tickseed

coreopsis, tick-seed

Habit Annuals, (5–)12–30(–50) cm. Annuals, perennials, subshrubs, or shrubs, 10–80(–200+) cm (often rhizomatous or with cormiform bases, stoloniferous in C. auriculata).
Stems

usually 1, erect, branched distally or ± throughout.

Leaves

lobes linear, 3–30 × 0.5–3 mm.

basal, basal and cauline, or cauline; opposite or alternate or both; petiolate or sessile;

blades simple and entire (dentate in C. latifolia), or ± pinnately or pedately lobed, faces glabrous or hairy.

Peduncles

5–25+ cm.

Involucres

± globose to cylindric, 4–25+ mm diam.

Receptacles

flat to convex, paleate;

paleae falling, ovate to linear or subulate, ± flat, ± scarious (not adnate to and not falling with cypselae except in C. bigelovii).

Ray florets

(5–)8+;

laminae 12–20(–30) mm.

mostly (5–)8(–12+, more in “double” cultivars), neuter, or styliferous and sterile, or pistillate and fertile;

corollas usually yellow, sometimes red-brown to purple proximally, sometimes wholly purple or pink to white.

Disc florets

(12–)40–100+;

corollas 4–6.6 mm.

8–150+, bisexual, fertile;

corollas usually yellow, sometimes red-brown to purple at tips or throughout, tubes equaling or shorter than narrowly funnelform throats, lobes 4 or 5 (adaxial sinus seldom deeper than others).

Phyllaries

(5–)8, lance-ovate to lanceolate, 6–10+ mm.

usually ± 8 in ± 2 series (usually distinct, rarely connate ± 1/10 their lengths, mostly oblong to linear, ± membranous, margins ± scarious).

Calyculi

of 3–5 ovate-deltate bractlets (3–)4–6 mm.

of (3–)8+, distinct, ± herbaceous bractlets.

Heads

radiate, borne singly or in open, ± corymbiform arrays.

Cypselae

obcompressed, ± orbiculate to ovate, oblong, or linear, usually thin-margined or winged, wings membranous to chartaceous or corky, entire or lobed to toothed, sometimes ciliolate;

faces smooth or ± papillate to tuberculate;

pappi 0, or persistent, of 2 bristly cusps or scales (sometimes pappi 0 and shoulders of cypsela wings ± bristly, pappus-like).

Disc cypselae

oblanceolate to linear, 5–6 mm;

pappi 3–4(–5) mm.

Ray cypselae

± oblong, 4–6 mm.

Paleae

seldom basally adnate to and falling with disc cypselae.

x

= 14.

2n

= 24.

Coreopsis calliopsidea

Coreopsis

Phenology Flowering Feb–Jun.
Habitat Alkaline playas, flood plains, deep sands
Elevation 100–1000 m (300–3300 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
from USDA
Mostly temperate North America; also tropical New World and Old World Early leaves of some coreopsises often differ from later leaves on individual plants
[BONAP county map]
Discussion

Species ca. 35 (28 in the flora).

Early leaves of some coreopsises often differ from later leaves on individual plants. In such plants, early (proximal) leaves may be 1–3-pinnately or pedately lobed and 12–25+ cm long with 9–15+ orbiculate to lanceolate, linear, or filiform lobes and contrast markedly with later (distal) leaves 1–3 cm that are undivided or have 3–5+ lanceolate to linear or filiform lobes.

Cultivars (often “doubles” with multiple series of “ray” florets) derived from Coreopsis auriculata, C. grandiflora, C. lanceolata, and C. tinctoria are grown in public and residential gardens and are grown commercially for cut flowers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ray florets usually pistillate and fertile, sometimes styliferous and sterile
→ 2
1. Ray florets neuter (without styles)
→ 4
2. Perennials, subshrubs, or shrubs; ray laminae ± elliptic to oblong, broadest at or near middles
sect. Tuckermannia
2. Annuals; ray laminae obovate to ± flabellate, broadest distal to middles
→ 3
3. Cypselae of disc florets ciliolate; pappi of 2 lanceolate to subulate scales
sect. Pugiopappus
3. Cypselae of disc florets not ciliolate; pappi 0 or coroniform (0.1–0.2 mm)
sect. Leptosyne
4. Leaf blades usually not lobed, rarely with 1–2 auricles at bases (margins dentate); rayflorets 4–5; disc florets 10–15+
sect. Silphidium
4. Leaf blades sometimes lobed (ultimate margins not dentate); ray florets (5–)8(–13; more in "double" cultivars); disc florets 30–150+
→ 5
5. Disc corolla lobes 5; style-branch apices usually ± penicillate, sometimes short-conic or truncate-mucronate
→ 6
5. Disc corolla lobes 4; style-branch apices ± truncate
→ 7
6. Ray laminae ± oblong-elliptic to suborbiculate, broadest at or near middles; paleae usually linear to subulate, often distally dilated
sect. Gyrophyllum
6. Ray laminae ± cuneate, broadest distal to middles; paleae basally ± lanceolate to linear, distally attenuate (± filiform)
sect. Coreopsis
7. Leaf blades sometimes with 1–2 lateral lobes; cypselae sometimes winged, wings ± pectinate or toothed
sect. Eublepharis
7. Leaf blades usually 1–2(–3)-pinnately lobed; cypselae sometimes winged, wingsentire
sect. Calliopsis
Source FNA vol. 21, p. 188. FNA vol. 21, p. 185. Author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Coreopsidinae > Coreopsis > sect. Pugiopappus Asteraceae > tribe Heliantheae > subtribe Coreopsidinae
Sibling taxa
C. auriculata, C. basalis, C. bigelovii, C. californica, C. delphiniifolia, C. douglasii, C. gigantea, C. gladiata, C. grandiflora, C. hamiltonii, C. integrifolia, C. intermedia, C. lanceolata, C. latifolia, C. leavenworthii, C. major, C. maritima, C. nudata, C. nuecensis, C. palmata, C. pubescens, C. pulchra, C. rosea, C. stillmanii, C. tinctoria, C. tripteris, C. verticillata
Subordinate taxa
C. sect. Calliopsis, C. sect. Coreopsis, C. sect. Eublepharis, C. sect. Gyrophyllum, C. sect. Leptosyne, C. sect. Pugiopappus, C. sect. Silphidium, C. sect. Tuckermannia
Synonyms Agarista calliopsidea
Name authority (de Candolle) A. Gray: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 90. (1859) Linnaeus: Sp. Pl. 2: 907. (1753): Gen. Pl. ed. 5, 388. (1754)
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