Cordylanthus capitatus |
Cordylanthus |
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Yakima bird's-beak, Yakima birdbeak |
bird's-beak, birdbeak |
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Habit | Herbs, annual; hemiparasitic. | |||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect or ascending, 10–50 cm, densely glandular-pubescent and pilose. |
erect or ascending, rarely decumbent, not fleshy, hairy or glabrous. |
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Leaves | glandular-pubescent and pilose; proximal 20–40(–50) mm, margins 3-lobed, lobes 1–2 mm wide; distal 10–30 × 1 mm, margins entire. |
cauline, alternate; petiole absent; blade not fleshy, not leathery, margins entire or 3–7-lobed. |
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Inflorescences | capitate spikes, 2–5-flowered; bracts 1–5, 10–20 mm, margins 3-lobed, lobes green and purple, linear to linear-lanceolate. |
terminal, spikes or flowers solitary, often capitate; bracts present. |
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Pedicels | bracteoles 12–18 mm, margins entire. |
absent; bracteoles absent. |
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Flowers | calyx 10–15 mm, tube 2–4 mm, apex 2-fid, cleft 3–5 mm; corolla purple-red, apex yellow, 10–20 mm, throat 4–6 mm diam., abaxial lip 3–5 mm, ca. equal to and appressed to adaxial; stamens 2, filaments glabrous, fertile pollen sacs 1 per filament, vestigial pollen sacs present. |
sepals 2, calyx bilaterally symmetric, spathelike, lobes triangular; petals 5, corolla yellow, yellow-green, purple, pink, or red, strongly bilabiate, club-shaped, abaxial lobes 3, middle lobe tightly revolute, tip distinctly folded inside-out, adaxial 2, adaxial lip galeate, rounded at apex, opening downward; stamens (2 or)4, didynamous, filaments hairy or glabrous; staminode 0; ovary 2-locular, placentation axile; stigma slightly expanded at apex. |
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Capsules | ovoid to oblong-ovoid, 5–7 mm. |
dehiscence loculicidal. |
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Seeds | 4–6, dark brown, narrowly ovoid to reniform, 2–2.5 mm, reticulate. |
4–25, pale brown to dark brown, ovoid to reniform, wings absent. |
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x | = 6, 7. |
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2n | = 26. |
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Cordylanthus capitatus |
Cordylanthus |
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Phenology | Flowering Jul–Oct. | |||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Openings in conifer woodlands and juniper scrub. | |||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 1400–2300 m. (4600–7500 ft.) | |||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; ID; NV; OR; WA
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w United States; nw Mexico |
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Discussion | Cordylanthus capitatus is often associated with Artemisia tridentata and may be hemiparasitic on it. Cordylanthus capitatus is distinguished by having only two anthers, each with one fertile pollen sac; infertile vestiges of the second pollen sac usually are present. The species can be recognized by its short-capitate inflorescences, the single, spathelike, deeply cleft calyx lobe, and the purple-red corollas with yellow tips. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 13 (13 in the flora). Cordylanthus is similar to Orthocarpus but differs in having spathelike calyces deeply cut along one side and narrowly lanceolate, entire bracteoles subtending each flower. Also, the abaxial lip of Orthocarpus is spreading; it usually is appressed to the adaxial lip in Cordylanthus. The mature flowers often appear to be buds. Cordylanthus is sometimes defined to include Dicranostegia and Chloropyron as subgenera (T. I. Chuang and L. R. Heckard 1986). D. C. Tank et al. (2009) found that those genera are a sister group of Triphysaria. The monograph by Chuang and Heckard provided the basis for most of the species delineation presented here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 671. | FNA vol. 17, p. 669. | ||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | C. nevadensis | Adenostegia | ||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Nuttall ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 597. (1846) | Nuttall ex Bentham: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 10: 597. (1846) — name conserved | ||||||||||||||||||||||||||||||||||||||||||||||||
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