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woolly sunflower

Habit Subshrubs, 50–150(–200) cm.
Stems

decumbent to ± erect, branched from bases or throughout (densely white-tomentose to glabrate).

Leaves

cauline; alternate;

petiolate;

blades broadly ovate, 1–2-pinnately lobed (lobes oblong or oblanceolate to linear), ultimate margins entire (somewhat revolute, apices ± rounded), faces white-tomentose (adaxial often glabrescent).

Involucres

cylindric to campanulate, 3–5 mm diam.

Receptacles

flat to convex, shallowly pitted, glabrous, epaleate.

Ray florets

4–9, pistillate, fertile;

corollas yellow.

Disc florets

10–25+, bisexual, fertile;

corollas yellow, tubes shorter than campanulate or narrowly funnelform throats, lobes 5, deltate.

Phyllaries

persistent, 8–16 in ± 2 series (± erect in fruit, distinct, oblong to linear, ± herbaceous, ± keeled).

Heads

radiate, (50–100+) in corymbiform or paniculiform arrays.

Cypselae

(blackish, dull) obpyramidal to clavate, ± scabrellous to glabrate;

pappi persistent, of 2–6+ unequal (or 2 opposite, longer, and ± equal), basally connate, oblong to subulate scales (tips acute to erose).

x

= 19.

Constancea

Distribution
from FNA
CA
[BONAP county map]
Discussion

Species 1.

Recognition of Constancea is based on morphologic, cytologic, and molecular evidence for polyphyly of Eriophyllum if E. nevinii is included. Constancea has characteristics found in other, closely related x = 19 “helenioid” genera that are unusual or absent in the clade represented by Eriophyllum, Pseudobahia, and Syntrichopappus, such as leaves with well-developed petioles, phyllaries in more than one series and more than the number of ray florets, and pappus scales unequal or a longer pair opposite and ± equal. The previously suggested close relationship between Constancea and another coastal subshrub, E. staechadifolium, is untenable.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 21, p. 362. Authors: Bruce G. Baldwin, John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Baeriinae
Subordinate taxa
C. nevinii
Name authority B. G. Baldwin: Madroño 46: 159. (2000)
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