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desert stinkweed, Hillman's cleomella, Hillman's stinkweed

stinkweed

Habit Plants 10–55(–85) cm. Herbs, usually annual [perennial] (usually unpleasantly scented).
Stems

sparsely branched proximally (central stem dominant); glabrous.

sparsely or profusely branched (usually erect); glabrous, pubescent, or scabrous.

Leaves

stipules 3–5-fid, 0.5–1 mm (scarious);

petiole 2–6(–8) cm;

leaflet blade elliptic, ovate, or ovate-oblong, 0.8–2.4 × 0.4–1.4 cm, thin, margins entire, apex obtuse to retuse, mucronulate, surfaces glabrous.

stipules (sometimes deciduous), 3–8-palmatifid, threadlike, or setaceous;

petiole without pulvinus (usually longer than leaflets);

leaflets (1 or) 3.

Inflorescences

racemes, terminating stems, 3–10 cm (2–20 cm in fruit);

bracts unifoliate, 5–9 mm.

terminal, racemes, corymbs, or flowers solitary in distal leaf axils (sometimes remaining compact in fruit);

bracts usually present (pedicels often anthocyanic).

Pedicels

divergent in fruit, 5–12 mm (4.5–17 mm in fruit).

Flowers

sepals green, ovate-lanceolate, 1–2 × 0.5–0.7 mm, glabrous;

petals yellow, oblong, (3.5–)4–8 × 1.8–2.2 mm, glabrous;

stamens (well-exserted), yellow, 8–12 mm;

anthers 2.5–3 mm;

gynophore ascending, 3.5–15 mm in fruit;

ovary ovoid, 1–1.3 mm;

style (0.7–)1–2 mm.

actinomorphic;

sepals persistent, connate basally (ca. 1/4 of length) [distinct], equal (each often subtending a nectary);

petals equal;

stamens 6 (usually exserted, distinct, equal);

filaments inserted on a discoid or conical androgynophore, glabrous;

anthers (ellipsoid), coiling as pollen is released;

gynophore reflexed, spreading, or ascending in fruit.

Fruits

capsules, dehiscent (as long as or shorter than wide), rhomboidal, globose, deltoid, or ovoid (valves laterally expanded, falling away from round, persistent septum).

Capsules

rhomboidal, 3.5–6 × 4–10.5 mm, glabrous.

Seeds

(1 or) 2–6, stramineous, globose, 2–2.7 mm, smooth.

1–16, globose or reniform, not arillate, (cleft fused between ends).

x

= 20.

Cleomella hillmanii

Cleomella

Distribution
from FNA
CA; ID; NV; OR; UT
[WildflowerSearch map]
[BONAP county map]
from USDA
United States; Mexico
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 10 (8 in the flora).

The two extraterritorial species of Cleomella are C. mexicana de Candolle and C. perennis H. H. Iltis, both known from central and northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pedicels (6.5-)9-17 mm in fruit; gynophore (6-) 7-15 mm in fruit; capsules 4-7(-8) mm diam.
var. hillmanii
1. Pedicels 4.5-8.5(-10) mm in fruit; gynophore 3.5-6.5 mm in fruit; capsules (5-)6.5-10.5 mm diam.
var. goodrichii
1. Gynophore 0.5-3 mm in fruit; petals 1.5-2.8 mm
→ 2
1. Gynophore 2.5-17 mm in fruit; petals 3-9 mm
→ 3
2. Pedicels recurved in fruit (± equal to gynophore), 1-2 mm; inflorescences solitary flowers in axils of closely spaced leaves.
C. brevipes
2. Pedicels spreading in fruit (much longer than gynophore), 12-20 mm (in fruit); inflorescences racemes, terminating stems and branches.
C. parviflora
3. Leaf blade surfaces (and sometimes stems) spreading-hispidulous; styles 2-3(-5) mm; plants usually spreading or matlike, sometimes erect.
C. obtusifolia
3. Leaf blade surfaces (and stems) glabrous; styles 0.3-2 mm; plants ± erect
→ 4
4. Gynophore 2.5-6(-8) mm in fruit; bracts ± rudimentary; plants without dominant central stem.
C. palmeriana
4. Gynophore 3.5-17 mm in fruit; bracts unifoliate or trifoliate (5-25 mm); plants with dominant central stem
→ 5
5. Leaflet blades ovate-elliptic, oblanceolate, or oblong-lanceolate, 0.4-1.4 cm wide (usually 1.5-2.5 times longer than wide)
→ 6
5. Leaflet blades linear to elliptic, 0.2-0.8 cm wide (usually 3-10 times longer than wide)
→ 7
6. Petioles 2-6(-8) cm; petals 1.8-2.2 mm wide; anthers 2.5-3 mm; California, Idaho, Nevada, Oregon, Utah.
C. hillmanii
6. Petioles 0.8-2 cm; petals 2-4 mm wide; anthers 1.5-1.9 mm; Arizona, New Mexico, Texas.
C. longipes
7. Sepals deltate, 0.5-1 mm wide; styles 0.3-0.5 mm; gynophore reflexed in fruit; capsules 4-6 × 6-12 mm, rhomboidal; seeds 2-3 mm, rugose.
C. angustifolia
7. Sepals lanceolate, 0.5-0.7 mm wide; styles 0.8-1.2 mm; gynophore ascending in fruit; capsules 2.5-5 × 2.5-6(-8) mm, ovoid to rhomboidal; seeds 2-2.2 mm, smooth.
C. plocasperma
Source FNA vol. 7, p. 210. FNA vol. 7, p. 209. Author: Staria S. Vanderpool.
Parent taxa Cleomaceae > Cleomella Cleomaceae
Sibling taxa
C. angustifolia, C. brevipes, C. longipes, C. obtusifolia, C. palmeriana, C. parviflora, C. plocasperma
Subordinate taxa
C. hillmanii var. goodrichii, C. hillmanii var. hillmanii
C. angustifolia, C. brevipes, C. hillmanii, C. longipes, C. obtusifolia, C. palmeriana, C. parviflora, C. plocasperma
Name authority A. Nelson: Proc. Biol. Soc. Wash. 18: 171. (1905) de Candolle: in A. P de Candolle and A. L. P. P. de Candolle, in A. Pde Candolle and A. L. P. P. de Candolle, Prodr. 1: 237. (1824)
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