Cleomella brevipes |
Cleomaceae |
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short-pedicelled cleomella, shortstalk cleomella, shortstalk stinkweed |
spider-flower family |
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Habit | Plants 3–15(–35) cm. | Herbs or shrubs, annual or perennial (usually deciduous, evergreen in Peritoma arborea); spines usually absent (present in Hemiscola and Tarenaya); glabrous or glandular-pubescent, hairs stalked or sessile (producing glucosinolates). | ||||||||||||||||||||||||||||||||||||||||||||||||
Stems | profusely branched (erect or spreading and matlike); glaucous, scabrous. |
usually erect, sometimes spreading or procumbent; branched or unbranched. |
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Leaves | stipules finely divided into crinkled threads, 2 mm; petiole 0.1–0.3 cm; leaflet blade linear-obovate, 0.5–1.5 × 0.1–0.3 cm, fleshy, margins entire, apex obtuse to acute, mucronate, surfaces scabrous. |
alternate, spirally arranged (usually palmately compound, sometimes simple); venation pinnate; stipules usually present (usually caducous, sometimes deciduous, 3–8-palmatifid, linear, threadlike, minute, scalelike, or absent, nodal (stipular) spines present in Tarenaya and Hemiscola); petiole present (pulvinus usually present, nectaries absent, petiolar spines sometimes present, petiolules present); blade margins entire, serrate, or serrulate. |
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Inflorescences | solitary flowers in axils of closely spaced leaves throughout; bracts unifoliate, 8–10 mm. |
terminal or axillary, usually racemose, sometimes flat-topped, or flowers solitary (usually elongated in fruit); bud scales absent; bracts present or absent (unifoliate, often trifoliate proximally, bracteoles absent). |
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Pedicels | reflexed in fruit, 1–2 mm. |
present. |
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Flowers | sepals green, ovate, 0.8–1.2 × 0.3–0.7 mm, glabrous; petals pale yellow, oblong, 1.5–2 × 0.5–1 mm, glabrous; stamens yellow, 1.5–2.2 mm; anthers 0.3–0.5 mm; gynophore strongly reflexed, 0.5–3 mm in fruit; ovary globose, 1–1.5 mm; style 2–3(–5) mm. |
usually bisexual (developmentally unisexual within sections of racemes), actinomorphic or slightly zygomorphic, rotate to crateriform, campanulate, or urceolate; perianth and androecium hypogynous; sepals persistent or deciduous, 4, distinct or connate basally; petals 4, attached directly to receptacle, imbricate, distinct, equal or unequal; intrastaminal nectary-discs, scales, or glands present or absent; stamens [4–]6–27[–35]; filaments free or basally adnate to gynophore (or along proximal 1/3–1/2 in Gynandropsis) or androgynophore, glabrous or pubescent; anthers dehiscing by longitudinal slits, pollen shed in single grains, binucleate, commonly tricolporate; gynophore present or absent; pistil 1; ovary 1-carpellate (except 2 in Oxystylis), 2-locular; placentation parietal; ovules 1–18(–26+) per locule, anatropous, bitegmic; style 1 (straight, relatively short, thick, not spinelike in fruit, except in Oxystylis, sometimes in Wislizenia); stigma 1, capitate, unlobed. |
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Fruits | capsular or nutlets (usually stipitate from elongation of gynophore, erect to divergent, usually not inflated), valvate, elongate (± dehiscent by 2 lateral valves, except in Polanisia), or schizocarps (inflated in Peritoma arborea), indehiscent or dehiscent. |
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Capsules | globose or deltoid, 2–3 × 2–3.2 mm, glabrous, (valves triangular). |
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Seeds | 1–4, yellowish, globose, 1.3–1.7 mm, smooth. |
1–65[–200], tan, yellowish brown, light brown, pale green, brown, reddish brown, silver-gray, or gray to black (papillose or tuberculate); arillate or not; endosperm scanty or absent, persistent perisperm sometimes present; cotyledons incumbent, (radicle-hypocotyl elongated). |
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Cleomella brevipes |
Cleomaceae |
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Phenology | Flowering spring–fall. | |||||||||||||||||||||||||||||||||||||||||||||||||
Habitat | Alkaline marshes, wet saline soils around thermal springs | |||||||||||||||||||||||||||||||||||||||||||||||||
Elevation | 400-1400 m (1300-4600 ft) | |||||||||||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; NV
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Nearly worldwide; tropical and temperate regions |
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Discussion | Genera 17, species ca. 150 (12 genera, 34 species in the flora). A discussion of the status of Cleomaceae and its segregation from Capparaceae (in the narrow sense) appears under the latter. Throughout this treatment, style length indicates the length in fruit; in some species, the style elongates after anthesis. The key to genera, in some cases, include characteristics of species, in addition to those of genera. This circumscription of Cleomaceae includes Oxystylidaceae Hutchinson. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 210. | FNA vol. 7, p. 199. | ||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | Cleomaceae > Cleomella | |||||||||||||||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | S. Watson: Proc. Amer. Acad. Arts 17: 365. (1882) | Berchtold & J. Presl | ||||||||||||||||||||||||||||||||||||||||||||||||
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