Clarkia stellata
Onagraceae tribe Onagreae
Lake Almanor clarkia, lake amador clarkia
erect, to 100 cm, puberulent.
petiole 5–30 mm;
blade lanceolate to elliptic or ovate, 1–5 cm.
alternate or basal;
stipules absent.
open racemes, axis in bud recurved 1–3 nodes distal to open flowers;
buds pendent, narrowly obovoid, tip acute.
floral tube 1.5–2 mm;
sepals reflexed individually;
corolla rotate, petals lavender-purple, not dark-flecked or spotted, obovate, 6–8 × 3–5 mm, inconspicuously 3-lobed;
stamens 8, subequal, subtended by ciliate scales, pollen yellow;
ovary shallowly 4-grooved, puberulent;
stigma not exserted beyond anthers.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
20–25 mm;
pedicel 1–3 mm.
unknown.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14.
Clarkia stellata
Onagraceae tribe Onagreae
Clarkia stellata is known from the southern Cascade–northern Sierra Nevada region, including Lassen, Nevada, Plumas, Shasta, Sierra, and Tehama counties (with unverified reports from Butte and Modoc counties).
Clarkia stellata is probably a self-pollinating derivative of C. mildrediae subsp. lutescens, to judge from pollen color. The two species are readily distinguishedby the much smaller flowers of C. stellata and the position of the stigma. Hybrids have low fertility due to chromosomal rearrangement. Clarkia stellata is morphologically very similar to the self-pollinating tetraploid C. rhomboidea but can be distinguished from it by yellow pollen and shallowly lobed, unspotted petals.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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